Syntaxonomical checklist of vascular plant communities of Spain and Portugal to association level

Salvador Rivas-Martínez, Federico Fernández-González, Javier Loidi, Mario Lousã & Angel Penas

Itinera Geobotanica 14: 5-341 (2001)


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Floating or rooted submerged aquatic vegetation

Grouped into nine major and twenty secondary categories, similar to the classic Braun-Blanquet's criterion of ecological progression, all the principal and supplementary ranks of syntaxa from class to association that we accept in Spain and Portugal (Iberian Peninsula, Balearic, Madeira, Azorean and Canary Islands) are listed alphabetically. In the present version, the Spanish and Portuguese hierarchical syntaxonomical checklist has 76 classes, 140 orders, 400 alliances and about 2000 associations. In all about 4000 syntaxa names (abstract units of phytocoenoses at any rank) we have aimed to comply with the definitions, principles, rules and recommendations of the 3rd edition of the International Code of Phytosociological Nomenclature [H.E. Weber, J. Moravec. & J.P. Théurillat in J. Veg. Sci. 11(5): 739-768.2000]. We have finally compiled around 1200 publications, only those ones where new syntaxa have been published.

I. FLOATING OR ROOTED SUBMERGED AQUATIC VEGETATION

Vegetación acuática flotante, sumergida o enraizada
Vegetação aquática flutuante, submersa ou enraizada

IA. FRESH-WATER VEGETATION

Vegetación de aguas dulces
Vegetação de água doce

1. Charetea fragilis Fukarek ex Krausch 1964

[Charetea fragilis Fukarek 1961 (art. 8)]

1a. Charetalia hispidae Sauer ex Krausch 1964

[Lamprothamnietalia papulosi van Raam & Schaminée in Schaminée, Weeda & Westhoff 1995 (art. 3b)]

1.1. Charion fragilis Krausch 1964

[Limno-Charion Krausch 1968 (art. 3a) p.p., Charion asperae Krause 1969 (art. 29), Charion contrario-asperae Pietsch 1987 (art. 29), Charion rudis-hispidae Pietsch 1987 (syntax. syn.), Thero-Charion asperae Krause 1969 (art. 22)]

1.1.1. Charetum asperae Corillion 1957

1.1.2. Charetum conniventis Velayos, Carrasco & Cirujano 1989

1.1.3. Charetum fragilis Corillion 1957

1.1.4. Charetum hispidae Margalef 1947

1.2. Charion vulgaris (Krause ex Krause & Lang 1977) Krause 1981

[Charenion vulgaris Krause 1969]

1.2.1. Charetum braunii Corillion 1957

1.2.2. Charetum imperfectae Cirujano 1995

1.2.3. Charetum vulgaris Corillion 1957

1.2.4. Tolypelletum glomeratae Corillion 1957

1.3. Charion canescentis Krausch 1964

[Halo-Charion Krausch 1968 (art. 3a, 29)]

1.3.1. Charetum canescentis Corillion 1957

1.3.2. Charetum galioidis Corillion 1957

1.3.3. Lamprothamnetum papulosi Corillion 1957

1.3.4. Tolypelletum hispanicae Corillion 1957

1b. Nitelletalia flexilis Krause 1969

1.4. Nitellion flexilis Dambska 1966 em. Krause 1969

[Nitellion Segal 1965 (art. 8), Nitellion flexilis Corillion ex Krause 1969 (art. 22)]

1.4.1. Nitelletum confervaceae Corillion 1957

1.4.2. Nitelletum flexilis Corillion 1957

1.4.3. Nitelletum opacae Corillion 1957

1.5. Nitellion syncarpo-tenuissimae Krause 1969

1.5.1. Tolypelletum proliferae Guerlesquin 1961

2. Lemnetea Tüxen ex O. Bolòs & Masclans 1955

[Lemnetea Tüxen 1953 (art. 8), Lemnetea Tüxen 1955 (art. 8), Lemnetea minoris Koch & Tüxen ex Schwabe & Tüxen 1981 (art. 31), Hydrocharitetea morsus-ranae Oberdorfer, Görs, Korneck, Lohmeyer, Müller, Philippi & Seibert 1967 (art. 2b, 8)]

2a. Lemnetalia minoris Tüxen ex O. Bolòs & Masclans 1955

[Lemnetalia Tüxen 1955 (art. 8), Lemnetalia minoris Tüxen ex Schwabe & Tüxen 1981 (art. 31)]

2.1. Lemnion minoris Tüxen ex O. Bolòs & Masclans 1955

[Lemnion minoris Tüxen 1955 (art. 8), Lemnion gibbae Tüxen & Schwabe in Tüxen 1974 (syntax. syn.), Lemno-Salvinion natantis Slavnic 1956 (syntax. syn.)]

2.1.1. Lemnetum gibbae Miyawaki & J. Tüxen 1960

2.1.2. Lemnetum minoris Oberdorfer ex Müller & Görs 1960

2.1.3. Lemno minoris-Salvinietum natantis (Slavnic 1956) Korneck 1959

2.1.4. Lemno-Azolletum filiculoidis Br.-Bl. in Br.-Bl., Roussine & Nègre 1952

[Lemno-Azolletum azolletosum carolinianae O. Bolòs & Masclans 1955 (corresp. name)]

2.1.5. Lemno-Spirodeletum polyrhizae Koch 1954

2.2. Lemnion trisulcae Den Hartog & Segal ex Tüxen & Schwabe in Tüxen 1974

[Riccio fluitantis-Lemnion trisulcae Schwabe & Tüxen 1981 (art. 29)]

2.2.1. Lemnetum trisulcae (Kelhofer 1915) Knapp & Stoffers 1962

2.2.2. Riccietum fluitantis Slavnic 1956 em. Tüxen 1974

2.2.3. Ricciocarpetum natantis Segal 1963 em. Tüxen 1974

2.3. Lemno minoris-Hydrocharition morsus-ranae Rivas-Martínez, Fernández-González & Loidi 1999

[Hydrocharition morsus-ranae Passarge 1996, non Rübel 1933]

2.3.1. Lemno minoris-Hydrocharitetum morsus-ranae Passarge 1978

3. Potametea Klika in Klika & Novák 1941

[Potametales Klika in Klika & Novák 1941 (original name), Potametea Tüxen & Preising 1942 (art. 8), Nymphaeetea Klika in Klika & Hadač 1944 (art. 29), Potametea Tüxen & Preising ex Oberdorfer 1957 (art. 31), Stratiotetea Den Hartog & Segal 1964 (syntax. syn.), Ceratophylletea Den Hartog & Segal 1964 (art. 8), Utricularietea neglectae Den Hartog & Segal 1964 (syntax. syn.)]

3a. Potametalia Koch 1926

[Hydrocharitetalia Rübel 1933 (syntax. syn.), Magnopotametalia Den Hartog & Segal 1964 (syntax. syn.), Parvopotametalia Den Hartog & Segal 1964 (art. 29), Callitricho-Ranunculetalia Passarge 1978 (syntax. syn.), Nymphaeetalia albo-tetragonae Passarge 1978 (syntax. syn.), Callitricho-Potametalia Schaminée, Lanjouw & Schipper 1990 (art. 29), Nupharo-Potametalia Schaminée, Lanjouw & Schipper 1990 (art. 29), Ranunculo-Myriophylletalia Passarge 1996 (syntax. syn.)]

3.1. Potamion (Koch 1926) Libbert 1931

[Potamion eurosibiricum Koch 1926 (art. 34), Parvo-Potamion eurosibiricum Vollmar 1947 (art. 34), Magno-Potamion eurosibiricum Vollmar 1947 (art. 34), Eu-Potamion (Koch 1926) Oberdorfer 1957 (art. 22), Magnopotamion (Vollmar 1947) Den Hartog & Segal 1964 (art. 22), Hydrocharition morsus-ranae Rübel 1933 (syntax. syn.), Stratiotion Den Hartog & Segal 1964 (syntax. syn.), Potamion pectinati (Koch 1926) Görs in Oberdorfer 1977 (art. 22)]

3.1.1. Groenlandio densae-Zannichellietum peltatae Velayos, Carrasco & Cirujano 1989

3.1.2. Myriophyllo alterniflori-Potametum crispi Rivas Goday 1964

3.1.3. Myriophyllo verticillati-Potametum pectinati Costa, Boira, Peris & Stübing 1986

3.1.4. Potametum colorati Allorge 1922

3.1.5. Potametum denso-nodosi O. Bolòs 1957

3.1.6. Potametum lucentis Hueck 1931

3.1.7. Potametum pectinati Cartensen 1955

3.1.8. Potametum perfoliato-crispi Bellot 1951

[ass. de Potamogeton perfoliatus y Potamogetum crispus Bellot 1951 (art. 14)]

3.1.9. Potametum trichoidis (Freitag, Markus & Schwipel 1958) Tüxen 1974

3.1.10. Potamo pectinati-Myriophylletum spicati Rivas Goday 1964 corr. Conesa 1990

3.1.11. Ranunculo trichophylli-Groenlandietum densae (Kohler, Brinkmeier & Vollrath 1974) Passarge 1994

3.1.12. Ranunculo eradicati-Potametum alpini Ballesteros & Gacia 1991

3.1.13. Stratiotetum aloidis Nowinski 1930

3.1.14. Zannichellio palustris-Potametum colorati O. Bolòs & Molinier in O. Bolòs 1996

3.2. Nymphaeion albae Oberdorfer 1957

3.2.1. Myriophyllo alterniflori-Potametum natantis ass. nova (addenda)

3.2.2. Nymphaeetum albo-luteae Nowinski 1928

[Myriophyllo verticillati-Nupharetum lutei Koch 1926 (art. 2b), Potamo-Vallisnerietum nymphaetosum albae Curcó 1996 (syntax. syn.)]

3.2.3. Limnanthemetum nymphoidis Bellot 1951

[ass. de Limnanthemum nymphoides Bellot 1951 (art. 14), Nymphoidetum peltatae Bellot 1951 nom. mut. (art. 45) (addenda)]

3.3. Ranunculion aquatilis Passarge 1964

[Callitricho-Batrachion Den Hartog & Segal 1964 (art. 22), Ranunculion peltati Schaminée, Lanjouw & Schipper 1990 (art. 29)]

3.3.1. Callitrichetum regis-jubae Galán in A.V. Pérez, Galán, P. Navas, D. Navas, Y. Gil & Cabezudo 1999

3.3.2. Callitricho platycarpae-Elatinetum alsinastri Rivas-Martínez & Costa in Rivas-Martínez, Costa, Castroviejo & E. Valdés 1980

3.3.3. Callitricho brutiae-Ranunculetum peltati ass. nova (addenda)

3.3.4. Callitricho stagnalis-Ranunculetum saniculifolii Galán in A.V. Pérez, Galán, P. Navas, D. Navas, Y. Gil & Cabezudo 1999

3.3.5. Myriophyllo alterniflori-Callitrichetum brutiae Cirujano, Pascual & Velayos 1986

3.3.6. Ranunculetum aquatilis (Sauer 1947) Géhu 1961

3.3.7. Ranunculetum baudotii Br.-Bl. in Br.-Bl., Roussine & Nègre 1952

[Callitricho-Ranunculetum aquatilis O. Bolòs, Molinier & P. Montserrat 1970, Callitricho-Ranunculetum baudotii O. Bolòs, Molinier & P. Montserrat 1970 corr. (art. 43) (addenda)]

3.3.8. Ranunculetum saniculifolii ass. nova (addenda)

3.3.9. Ranunculetum tripartiti Galán in A.V. Pérez, Galán, P. Navas, D. Navas, Y. Gil & Cabezudo 1999

3.3.10. Ranunculo tripartiti-Myriophylletum alterniflori Franquesa 1995

3.4. Ranunculion fluitantis Neuhäusl 1959

3.4.1. Callitricho brutiae-Ranunculetum pseudofluitantis ass. nova (addenda)

3.4.2. Callitricho lusitanicae-Ranunculetum penicillati ass. nova (addenda)

[Callitricho lusitanicae-Ranunculetum penicillati Pizarro 1995 (art. 3b)]

3.4.3. Ranunculetum fluitantis Allorge 1922

3.5. Zannichellion pedicellatae Schaminée, Lanjouw & Schipper 1990 em. Pott 1992

3.5.1. Najadetum marinae Fukarek 1961

[Parvopotamo-Najadetum marinae Oberdorfer 1957 (art. 2b), Parvopotamo-Najadetum marinae Oberdorfer ex Knapp & Sell 1965 (syntax. syn.)]

3.5.2. Zannichellietum pedicellatae Nordhagen 1954

3b. Utricularietalia Den Hartog & Segal 1964

[Lemno-Utricularietalia vulgaris Passarge 1978 (syntax. syn.)]

3.6. Utricularion Den Hartog & Segal 1964

[Utricularion vulgaris Passarge (1964) 1996 (syntax. syn.)]

3.6.1. Lemno minoris-Utricularietum vulgaris (Soó 1928) Passarge 1964

3.6.2. Potamo-Utricularietum australis Br.-Bl. in Br.-Bl., Roussine & Nègre 1952 corr. (addenda)

[Potamo-Utricularietum vulgaris Br.-Bl. in Br.-Bl., Roussine & Nègre 1952 (art. 43)]

3.6.3. Utricularietum exoleto-australis Rivas-Martínez, Costa, Castroviejo & E. Valdés 1980

3.7. Ceratophyllion demersi Den Hartog & Segal ex Passarge 1996

[Ceratophyllion demersi Den Hartog & Segal 1964 (art. 8)]

3.7.1. Potamo-Ceratophylletum demersi Hild & Rehnelt 1965

3.7.2. Potamo-Ceratophylletum submersi Pop 1962

[Ceratophylletum submersi Den Hartog & Segal ex Redeker 1969]

IB. MARINE AND SALT-WATER VEGETATION

Vegetación marina y de aguas saladas
Vegetação marinha e de água salgada

4. Halodulo wrightii-Thalassietea testudinum Den Hartog ex Rivas-Martínez, Fernández-González & Loidi 1999

[Halodulo-Thalassietea Den Hartog 1976 (art. 8), Halodulo-Cymodoceetea Knapp 1965 (art. 8)]

4a. Thalassio-Syringodietalia filiformis Borhidi, Muñiz & Del Risco in Borhidi 1996

[Thalassietalia Den Hartog 1976 (art. 8), Thalassio-Syringodietalia filiformis Borhidi, Muñiz & Del Risco 1983 (art. 5)]

4.1. Syringodio-Thalassion testudinum Borhidi 1996

[Cymodoceion nodosae Den Hartog 1976 (art. 8)]

4.1.1. Cymodoceetum nodosae Feldmann 1937

4.1.2. Halophiletum decipientis ass. nova (addenda)

5. Posidonietea Den Hartog 1976 (art. 8)

5a. Posidonietalia Den Hartog 1976

5.1. Posidonion Br.-Bl., Roussine & Nègre 1952

[Posidonion Br.-Bl. 1931 (art. 8)]

5.1.1. Posidonietum oceanicae Funk 1927

6Ruppietea J. Tüxen 1960

[Rielletea helicophyllae Cirujano, Velayos & P. García 1993 (art. 5, 8)]

6a. Ruppietalia maritimae J. Tüxen 1960

[Zannichellietalia pedicellatae Schaminée, Lanjouw & Schipper 1990 (syntax. syn.), Zannichellio-Ruppietalia Schaminée & Den Hartog in Schaminée, Weeda & Westhoff 1995 (art. 29), Rielletalia helicophyllae Cirujano, Velayos & P. García 1993 (art. 5, 8)]

6.1. Ruppion maritimae Br.-Bl. ex Westhoff in Bennema, Sissingh & Westhoff 1943

[Ruppion maritimae Br.-Bl. 1931 (art. 8), Althenio-Ruppion Den Hartog 1981 (art. 8)]

6.1.1. Ruppietum spiralis Hocquette 1927 corr. Iversen 1934

[ass. Ruppia maritima Hocquette 1927 (art. 43), Chaetomorpho limi-Ruppietum Br.-Bl. in Br.-Bl., Roussine & Nègre 1952 (syntax. syn.), Ruppietum cirrhosae Hocquette 1927 corr. Iversen 1934 nom. mut. (art. 45) (addenda)]

6.1.2. Ruppietum drepanensis Brullo & Furnari 1976

6.1.3. Enteromorpho intestinalidis-Ruppietum maritimae Westhoff ex Tüxen & Böckelmann 1957

[Ruppietum maritimae Iversen 1934 (art. 31), Enteromorpha intestinalis-Ruppia rostellata ass. Westhoff in Bennema, Sissingh and Westhoff 1943 (art. 1)]

6.2. Scirpion parvuli Segal 1968

6.2.1. Eleocharitetum parvulae (Christiansen 1934) Gillner 1960

6.3Riellion helicophyllae Cirujano, Velayos & P. García ex Rivas-Martínez, Fernández-González & Loidi 1999

[Riellion helicophyllae Cirujano, Velayos & P. García 1993 (art. 5, 8)]

6.3.1. Rielletum helicophyllae Cirujano, Velayos & P. García 1993

6.3.2. Rielletum notarisii Cirujano, Velayos & P. García 1993

7. Zosteretea marinae Pignatti 1954

7a. Zosteretalia Béguinot 1941

7.1. Zosterion Christiansen 1934

7.1.1. Guiraudio-Zosteretum noltii Boudouresque, Belscher & Marcot-Coqueugniot 1977

7.1.2. Zosteretum marinae (Borgesen 1905) Harmsen 1936

7.1.3. Zosteretum noltii Harmsen 1936