Vascular plant communities of Spain and Portugal. Addenda to the Syntaxonomical checklist of 2001

Salvador Rivas-Martínez, Tomás E. Díaz, Federico Fernández-González, Jesús Izco, Javier Loidi, Mario Lousã & Ángel Penas

Itinera Geobotanica 15(1-2): 5-922 (2002)


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Syntaxonomical system: Coastal and continental halophilous and sand dune vegetation

IIIa. Vegetation of coastal sand dunes

16. Ammophiletea Br.-Bl. & Tüxen ex Westhoff, Dijk & Passchier 1946

Pioneer perennial rhizomatous grasses and chamaephyte coastal sand dune communities of Mediterranean and Atlantic littoral distribution. Mobile fore-dunes (Ammophiletalia) [external flat coastal dunes (Honckenyo-Elytrigion: Elytrigienion boreoatlanticae = Atlantic, Coastal Lusitan-Andalusian and Tingitanian, Elytrigienion junceae = Mediterranean excepting Tingitanian and Coastal Lusitan-Andalusian), internal high coastal dunes (Ammophilion australis: Ammophilenion australis = Mediterranean and Cantabrian-Atlantic until Basque Country coasts), Mediterranean salt flat coastal dunes (Sporobolion arenarii)]; grey hind-dunes (Crucianelletalia maritimae) [Mediterranean drainage basin (Crucianellion maritimae), franco-atlantic coasts (Euphorbio-Helichrysion maritimae), ibero-atlantic coasts (Helichrysion picardii)]. Mediterranean and Atlantic-Central European.
Characteristic species: Anthemis maritima, Cyperus capitatus, Euphorbia paralias, Lotus creticus, Medicago marina, Pancratium maritimum, Polygonum maritimum.

16a. Ammophiletalia Br.-Bl. 1933
Characteristic species: Calystegia soldanella, Eryngium maritimum, Festuca juncifolia, Otanthus maritimus.

16.1. Ammophilion australis Br.-Bl. 1921 corr. Rivas-Martínez, Costa & Izco in Rivas-Martínez, Lousã, T.E. Díaz, Fernández-González & J.C. Costa 1990
[Ammophilion arundinaceae]
( Ammophilenion australis Rivas-Martínez & Géhu in Rivas-Martínez, Costa, Castroviejo & E. Valdés 1980 corr. F. Prieto & T. E. Díaz 1991 (16.1a) )
Characteristic species: Ammophila arenaria subsp. australis (16.1a), Echinophora spinosa (16.1a).

16.2. Honckenyo peploidis-Elytrigion boreoatlanticae Tüxen in Br.-Bl. & Tüxen 1952 nom. mut. & inv. propos.
[Agropyro-Minuartion peploidis]
( Elytrigienion boreoatlanticae Rivas-Martínez & Géhu in Rivas-Martínez, Costa, Castroviejo & E. Valdés 1980 nom. mut. propos. (16.2a). Elytrigienion junceae Rivas-Martínez, Costa, Castroviejo & E. Valdés 1980 nom. mut. propos. (16.2b) )
Characteristic species: Elytrigia juncea subsp. boreoatlantica (16.2a), Elytrigia juncea subsp. juncea (16.2b), Honckenya peploides var. peploides (16.2a).

16.3. Sporobolion arenarii (Géhu & Géhu-Franck ex Géhu & Biondi 1994) Rivas-Martínez & Cantó all. nova, stat. nov.
Characteristic species: Centaurea seridis var. maritima, Centaurea sphaerocephala var. polyacantha, Sporobolus arenarius.

16b. Crucianelletalia maritimae Sissingh 1974
Characteristic species: Aetheorhiza bulbosa, Crucianella maritima, Euphorbia portlandica, Helichrysum stoechas var. maritimum, Leontodon taraxacoides subsp. taraxacoides, Malcolmia littorea, Matthiola sinuata, Ononis ramosissima, Scrophularia frutescens.

16.4. Crucianellion maritimae Rivas Goday & Rivas-Martínez 1958
Characteristic species: Ambrosia maritima, Echium sabulicola, Launaea fragilis subsp. viminea, Teucrium dunense.

16.5. Euphorbio portlandicae-Helichrysion maritimi Géhu & Tüxen ex Sissingh 1974
Characteristic species: Alyssum loiseleurii, Asperula occidentalis, Dianthus hyssopifolius subsp. gallicus, Galium arenarium, Linaria maritima.

16.6. Helichrysion picardii (Rivas-Martínez, Costa & Izco in Rivas-Martínez, Lousã, T.E. Díaz, Fernández-González & J.C. Costa 1990) Rivas-Martínez, Fernández-González & Loidi 1999
Characteristic species: Armeria pungens, Armeria welwitschii, Artemisia crithmifolia, Coincya pseudoerucastrum subsp. johnstonii, Echium gaditanum, Euphorbia baetica, Helichrysum picardii, Iberis procumbens, Jasione lusitanica, Linaria navarroi, Linaria lamarckii, Thymus carnosus.

17. Cakiletea maritimae Tüxen & Preising ex Br.-Bl. & Tüxen 1952

Pioneer halo-nitrophilous annual forb communities of strandlines, beaches and coastal sand dunes (Cakiletalia integrifoliae) [Atlantic and Baltic See coasts (Atriplicion littoralis), Mediterranean and Coastal Lusitan-Andalusian (Cakilion maritimae)]. Holarctic.
Characteristic species: Beta maritima var. maritima, Cakile edentula, Chamaesyce peplis, Glaucium flavum, Salsola kali subsp. kali, Salsola kali var. polysarca.

17a. Cakiletalia integrifoliae Tüxen ex Oberdorfer 1949 corr. Rivas-Martínez, Costa & Loidi 1992
[Cakiletalia maritimae]
Characteristic species: See class (17).

17.1. Atriplicion littoralis Nordhagen 1940
Characteristic species: Atriplex glabriuscula, Atriplex laciniata, Cakile maritima subsp. integrifolia, Chamaesyce polygonifolia, Polygonum raii (*), Raphanus maritimus subsp. maritimus.

17.2. Cakilion maritimae Pignatti 1953
Characteristic species: Atriplex rosea subsp. tarraconensis, Cakile maritima subsp. maritima.

18. Honckenyo-Leymetea arenarii Tüxen 1966 nom. mut. propos.

[Honckenyo-Elymetea arenarii]
European class not found in the territory.

IIIb. Coastal and continental halophilous vegetation

19. Crithmo-Limonietea Br.-Bl. in Br.-Bl., Roussine & Nègre 1952 nom. mut. propos.

[Crithmo-Staticetea]
Coastal chasmophytic communities of cliffs and lithosols splashed by marine salt spray under mediterranean bioclimate (Crithmo-Limonietalia) [Western Mediterranean (Crithmo-Limonion), Coastal Lusitan-Andalusian (Crithmo-Daucion halophili), Canarian (Frankenio-Astydamion) and Madeiran (Helichrysion obconico-devium) and dwarf chamaephyte cushion-shaped communities of lithosols or sandy-lithosols of mediterranean coastal rockies splashed by strong marine winds in Balearic Islands (Launaeion cervicornis) or locally in Iberian Peninsula (Sagres) and Catalan-Valencian (Astragalion tragacanthae)]. Mediterranean.
Characteristic species: Asteriscus maritimus, Crithmum maritimum, Limonium virgatum, Plantago macrorhiza.

19a. Crithmo-Limonietalia Molinier 1934 nom. mut. propos.
[Crithmo-Staticetalia]
Characteristic species: Helichrysum stoechas subsp. caespitosum.

19.1. Crithmo-Limonion Molinier 1934 nom. mut. propos.
[Crithmo-Staticion]
Characteristic species: Daucus gingidium subsp. commutatus, Daucus gingidium subsp. hispanicus, Euphorbia margalidiana, Frankenia laevis var. marcosii, Limonium artruchium, Limonium balearicum, Limonium caprariense, Limonium carregadorense, Limonium dragonericum, Limonium dufourii, Limonium ebusitanum, Limonium fontqueri, Limonium geronense, Limonium gymnesicum, Limonium majoricum, Limonium malacitanum, Limonium minoricense, Limonium minutum, Limonium muradense, Limonium portopetranum, Limonium pseudebusitanum, Limonium pseudodictyocladum, Limonium revolutum, Limonium saxicola, Limonium tenuicaule, Limonium tremolsii, Lotus cytisoides, Senecio leucanthemifolius subsp. crassifolius, Sonchus tenerrimus var. pectinatus.

19.2. Launaeion cervicornis (O. Bolòs & Vigo ex Gil & Llorens 1995) Rivas-Martínez, Fernández-González & Loidi 1999
Characteristic species: Anthyllis hystrix, Dorycnium fulgurans, Euphorbia maresii subsp. maresii, Femeniasia balearica, Launaea cervicornis, Santolina chamaecyparissus subsp. magonica.

19.3. Crithmo-Daucion halophili Rivas-Martínez, Lousã, T.E. Díaz, Fernández-González & J.C. Costa 1990
Characteristic species: Armeria pseudarmeria, Daucus halophilus, Dianthus cintranus subsp. cintranus, Limonium emarginatum, Limonium multiflorum, Limonium plurisquamatum, Silene obtusifolia, Spergularia australis.

19.4. Frankenio-Astydamion latifoliae Santos 1976
Characteristic species: Argyranthemum frutescens subsp. succulentum, Astydamia latifolia, Atractylis preuxiana, Frankenia ericifolia, Limonium humboldtii, Limonium imbricatum, Limonium papillatum, Limonium pectinatum, Reichardia crystalina, Reichardia ligulata var. crispa.

19.5. Astragalion tragacanthae (Folch ex Rivas-Martínez, Fernández-González & Loidi 1999) Rivas-Martínez, Fernández-González & Loidi all. nova, stat. nov.
Characteristic species: Astragalus tragacantha subsp. tragacantha, Astragalus tragacantha subsp. vicentinus, Festuca ruscinonensis, Plantago subulata, Polycarpon polycarpoides subsp. catalaunicum, Senecio cineraria, Seseli farrenyi, Silene rothmaleri.

19.6. Helichrysion obconico-devium Rivas-Martínez, Capelo, J.C. Costa, Lousã, Fontinha, Jardim & Sequeira all. nova
Characteristic species: Helichrysum devium, Helichrysum obconicum, Limonium pyramidatum.

20. Juncetea maritimi Br.-Bl. in Br.-Bl., Roussine & Nègre 1952

Perennial grasslands growing on coastal and inland temporary wet or inundated salt marshes. Temperate continental and low littoral Cantabrian-Atlantic (Glauco-Puccinellietalia) [coastal dense juncoid wet salt marshes (Glauco maritimae-Juncion maritimi), coastal prostratae chamaephyte halo-nitrophilous communities reached by highest tides (Limonio ovalifolii-Frankenion laevis)]; Mediterranean (Juncetalia maritimi) [coastal or inland Juncus maritimus and graminoid wet communities (Juncion maritimi: Juncenion maritimi = coastal euoceanic, Soncho crassifolii-Juncenion maritimi = inland semicontinental), coastal halophilous well drained sandy soils (Plantaginion crassifoliae), inland long stagnant brackish water (Puccinellion caespitosae)]; aerohaline Atlantic sea cliffs (Crithmo-Armerietalia) [Britanic, Cantabrian-Atlantic, Coastal Lusitan-Andalusian and West Tingitanian (Crithmo-Armerion), Azorean (Euphorbio azoricae-Festucion petraeae)]. Holarctic.
Characteristic species: Alopecurus bulbosus, Apium graveolens, Aster tripolium, Carex extensa, Festuca rubra subsp. pruinosa, Glaux maritima, Juncus gerardi, Juncus maritimus, Limonium auriculae-ursifolium, Plantago maritima, Puccinellia rupestris, Triglochin maritima.

20a. Juncetalia maritimi Br.-Bl. ex Horvatic 1934
Characteristic species: Dorycnium gracile, Elytrigia elongata, Juncus acutus, Lotus preslii, Melilotus siculus, Puccinellia festuciformis subsp. lagascana, Puccinellia tenuifolia, Sonchus maritimus, Spartina versicolor, Thalictrum maritimum.

20.1. Juncion maritimi Br.-Bl. ex Horvatic 1934
(Juncenion maritimi Rivas-Martínez 1984 (20.1a). Soncho crassifolii-Juncenion maritimi Rivas-Martínez 1984 (20.1b))
Characteristic species: Centaurium tenuiflorum, Elytrigia curvifolia (20.1b), Gypsophila tomentosa (20.1b), Helianthemum polygonoides (20.1b), Iris spuria subsp. maritima, Juncus subulatus, Sonchus crassifolius (20.1b), Tetragonolobus maritimus var. maritimus.

20.2. Plantaginion crassifoliae Br.-Bl. in Br.-Bl., Roussine & Nègre 1952
Characteristic species: Centaurea dracunculifolia, Linum maritimum, Plantago crassifolia.

20.3. Puccinellion caespitosae Rivas-Martínez in Rivas-Martínez & Costa 1976 corr. hoc loco
[Puccinellion fasciculatae]
Characteristic species: Carex punctata subsp. lainzii, Puccinellia caespitosa, Puccinellia hispanica, Puccinellia pungens, Scorzonera parviflora.

20b. Glauco-Puccinellietalia Beeftink & Westhoff in Beeftink 1962

Characteristic species: Armeria maritima subsp. maritima, Limonium binervosum, Limonium vulgare, Puccinellia distans, Puccinellia fasciculata, Puccinellia maritima.

20.4. Glauco maritimae-Juncion maritimi Géhu & Géhu-Franck 1984
Characteristic species: Carex punctata subsp. punctata, Cochlearia aestuaria, Festuca rubra subsp. litoralis, Limonium humile.

20.5. Limonio ovalifolii-Frankenion laevis Arbesú, Bueno & F. Prieto all. nova
Characteristic species: Frankenia laevis var. laevis, Limonium dodartii, Limonium ovalifolium.

20c. Crithmo-Armerietalia Géhu 1964
Characteristic species: Anthyllis vulneraria subsp. iberica, Dactylis marina.

20.6. Crithmo-Armerion maritimae Géhu 1968
Characteristic species: Angelica pachycarpa, Armeria berlengensis, Armeria euscadiensis, Armeria maritima subsp. miscella, Armeria pubigera subsp. depilata, Armeria pubigera subsp. pubigera, Centaurea borjae, Daucus gingidium subsp. atlanticum, Leucanthemum ircutianum subsp. crassifolium, Leucanthemum merinoi, Leucanthemum pluriflorum, Rumex acetosa subsp. biformis, Rumex rupestris, Sesamoides suffruticosa subsp. latifolia, Silene uniflora, Spergularia rupicola subsp. rupicola, Trifolium occidentale.

20.7. Euphorbio azoricae-Festucion petraeae Lüpnitz 1976
Characteristic species: Azorina vidali, Daucus azoricus, Euphorbia azorica, Festuca petraea, Spergularia azorica.

21. Puccinellio-Salicornietea Topa 1939

European class not found in the territory.

22. Saginetea maritimae Westhoff, Van Leeuwen & Adriani 1962

Pioneer ephemeral coastal and inland communities on bare or disturbed salt-marsh. Littoral Atlantic temperate, saline, and loamy or sandy soils under salt-spray influence (Saginetalia maritimae: Saginion maritimae); Mediterranean littoral or inland halophilous habitats (Frankenietalia pulverulentae) [coastal and inland long stagnant saline waters (Frankenion pulverulentae), inland slightly inundated saline waters (Hordeion marini)]. Holarctic.
Characteristic species: Bupleurum tenuissimum, Catapodium marinum, Centaurium spicatum, Parapholis incurva, Parapholis strigosa, Plantago coronopus var. stricta, Rhynchostegiella littorea, Sagina maritima, Spergularia bocconei, Spergularia marina.

22a. Saginetalia maritimae Westhoff, Van Leeuwen & Adriani 1962
Characteristic species: See alliance (22.1).

22.1. Saginion maritimae Westhoff, Van Leeuwen & Adriani 1962
Characteristic species: Amblystegium serpens var. salinum, Cochlearia danica, Sagina nodosa.

22b. Frankenietalia pulverulentae Rivas-Martínez ex Castroviejo & Porta 1976
Characteristic species: Frankenia pulverulenta, Parapholis filiformis, Parapholis pycnantha, Spergularia diandra.

22.2. Frankenion pulverulentae Rivas-Martínez ex Castroviejo & Porta 1976
Characteristic species: Arenaria viridis, Catapodium rigidum subsp. spicatum, Filago mareotica, Hymenolobus pauciflorus, Hymenolobus procumbens, Silene sedoides, Sphenopus divaricatus, Teucrium campanulatum.

22.3. Hordeion marini Ladero, F. Navarro, C. Valle, B. Marcos, Ruiz & M.T. Santos 1984
Characteristic species: Agrostis nebulosa, Gaudinia hispanica, Hainardia cylindrica, Hainardiopholis xpauneroi, Hordeum marinum, Parapholis pannonica, Polypogon maritimus subsp. maritimus, Polypogon maritimus subsp. subspathaceus, Rostraria phleoides, Rostraria pumila.

23. Sarcocornietea fruticosae Br.-Bl. & Tüxen ex A. & O. Bolòs 1950 nom. mut. propos.

[Salicornietea fruticosae]
Coastal and inland Mediterranean and Cantabrian-Atlantic perennial salt-marshes and salt-pans communities. Temporarily inundated succulent chenopodiaceous dwarf shrubs (Sarcocornietalia fruticosae) [coastal Mediterranean and Cantabrian-Atlantic to South England salt-marshes reached by tides or in saltworks (Sarcocornion fruticosae: Sarcocornienion fruticosae = Mediterranean and Cantabrian-Atlantic to South Britanny saltworks and wet salt-marshes occasionally reached by tides, Sarcocornienion perennis = Cantabrian-Atlantic to South England tidelands), coastal Mediterranean to South Galicia and related inland salt pans short time inundated (Arthrocnemion macrostachyi: Arthrocnemenion macrostachyi = Mediterranean and Coastal Lusitan-Andalusian to Lisbonne in high elevations of salt-marshes ever out of tides and related inland salt pans, Sarcocornienion alpini = Mediterranean, Coastal Lusitan-Andalusian and Cantabrian-Atlantic to South Galicia salt-marshes only exceptionally reached by tides), inland salt-pans winter inundated (Suaedion braun-blanquetii), coastal halo-nitrophilous slightly inundated (Suaedion verae)]; littoral and inland temporary wet Mediterranean high saline Limonium and perennial grasses (Limonietalia) [coastal Western Mediterranean dry and subhumid short time inundated Limonium communities (Limonion confusi), coastal thermomediterranean well drained sandy Limoniastrum monopetalum communities (Limoniastrion monopetali), inland mesomediterranean semiarid and dry Lygeum-Limonium communities of Ebro river drainage basin (Limonion catalaunico-viciosoi), inland mesomediterranean dry Lygeum-Limonium communities of Castilian biogeographic Subprovince (Lygeo-Lepidion cardamines), coastal and inland Murcian-Almeriensian thermo-mesomediterranean semiarid Lygeum-Limonium communities (Lygeo-Limonion angustebracteati)]. Holarctic.
Characteristic species: Artemisia gallica subsp. gallica, Inula crithmoides.

23a. Sarcocornietalia fruticosae Br.-Bl. 1933 nom. mut. propos.
[Salicornietalia fruticosae]
Characteristic species: Halimione portulacoides, Puccinellia festuciformis subsp. festuciformis, Triglochin barrelieri.

23.1. Sarcocornion fruticosae Br.-Bl. 1933 nom. mut. propos.
[Salicornion fruticosae]
(Sarcocornienion fruticosae Rivas-Martínez & Costa 1984 nom. mut. propos. (23.1a). Sarcocornienion perennis Rivas-Martínez in Rivas-Martínez & Costa 1984 nom. mut. propos. (23.1b))
Characteristic species: Sarcocornia fruticosa, Sarcocornia perennis (23.1b).

23.2. Arthrocnemion macrostachyi Rivas-Martínez & Costa 1984 nom. mut. propos.
[Arthrocnemion glauci]
(Arthrocnemenion macrostachyi Rivas-Martínez, Lousã, T.E. Díaz, Fernández-González & J.C. Costa 1990 nom. mut. propos. (23.2a). Sarcocornienion alpini Rivas-Martínez, Lousã, T.E. Díaz, Fernández-González & J.C. Costa 1990 (23.2b))
Characteristic species: Arthrocnemum macrostachyum, Frankenia corymbosa, Halocnemum strobilaceum, Limonium algarvense, Limonium daveaui, Limonium ferulaceum, Sarcocornia alpini (23.2b).

23.3. Suaedion braun-blanquetii Br.-Bl. & O. Bolòs 1958 corr. Rivas-Martínez, Báscones, T.E. Díaz, Fernández-González & Loidi 1991
[Suaedion brevifoliae]
Characteristic species: Suaeda braun-blanquetii.

23.4. Suaedion verae (Rivas-Martínez, Lousã, T.E. Díaz, Fernández-González & J.C. Costa 1990) Rivas-Martínez, Fernández-González & Loidi 1999
[Suaedenion verae]
Characteristic species: Suaeda vera.

23b. Limonietalia Br.-Bl. & O. Bolòs 1958
Characteristic species: Limonium bellidifolium, Limonium cossonianum, Limonium densissimum, Limonium latebracteatum, Limonium narbonense, Limonium supinum.

23.5. Limonion confusi (Br.-Bl. 1933) Rivas-Martínez & Costa 1984
Characteristic species: Limonium antonii-llorensii, Limonium biflorum, Limonium boirae, Limonium cavanillesii, Limonium companyonis, Limonium confusum, Limonium diffusum, Limonium gibertii, Limonium girardianum, Limonium grosii, Limonium heterospicatum, Limonium legrandii, Limonium magallufianum, Limonium marisolii, Limonium migjornense, Limonium pseudarticulatum, Limonium retusum, Limonium scorpioides, Limonium wiedmannii.

23.6. Limoniastrion monopetali Pignatti 1953
Characteristic species: Limoniastrum monopetalum.

23.7. Limonion catalaunico-viciosoi Rivas-Martínez & Costa 1984
Characteristic species: Artemisia gallica subsp. gargantae, Limonium aragonense, Limonium catalaunicum, Limonium costae, Limonium hibericum, Limonium ruizii, Limonium stenophyllum, Limonium viciosoi, Senecio auricula subsp. sicoricus.

23.8. Lygeo-Lepidion cardamines Rivas Goday & Rivas-Martínez ex Rivas-Martínez & Costa 1984
Characteristic species: Limonium carpetanicum, Limonium cofrentanum, Limonium dichotomum, Limonium erectum, Limonium lobetanicum, Limonium majus, Limonium minus, Limonium pinillense, Limonium quesadense, Limonium soboliferum, Limonium squarrosum, Limonium sucronicum, Limonium toletanum, Limonium tournefortii, Senecio auricula subsp. castellanus.

23.9. Lygeo-Limonion angustebracteati Rigual ex Alcaraz, P. Sánchez & De la Torre 1989
Characteristic species: Limonium angustebracteatum, Limonium caesium, Limonium cordovillense, Limonium delicatulum, Limonium furfuraceum, Limonium insigne, Limonium santapolense, Senecio auricula subsp. auricula.

24. Spartinetea maritimae Tüxen in Beeftink & Géhu 1973

Pioneer coastal Spartina perennial grass communities on intertidal mud and sand (Spartinetalia maritimae: Spartinion maritimae). Holarctic.
Characteristic species: Spartina alterniflora, Spartina densiflora, Spartina xtownsendii.

24a. Spartinetalia maritimae Conard ex Beeftink & Géhu 1973
Characteristic species: See alliance (24.1).

24.1. Spartinion maritimae Conard ex Beeftink & Géhu 1973
Characteristic species: Spartina maritima.

25. Thero-Salicornietea Tüxen in Tüxen & Oberdorfer ex Géhu & Géhu-Frank 1984 nom. conserv. propos.

[Thero-Suaedetea Rivas-Martínez 1972 (art. 52)]
Pioneer annual coastal and continental tidal mud-flat and temporary inundated salt marshes and salt-pans communities of glassworts and other therophyte succulent plants. Littoral and inland halophytic (Thero-Salicornietalia) [tidal muddy flats mostly Cantabrian-Atlantic tetraploids (Salicornion dolichostachyo-fragilis), upper muddy flats and salt marsh coastal and continental Atlantic-Central European diploids (Salicornion europaeo-ramosissimae), Mediterranean coastal and inland long inundated diploids and tetraploids (Salicornion patulae), Mediterranean continental temporary wet (Microcnemion coralloidis)]; littoral and inland halo-nitrophilous (Thero-Suaedetalia: Thero-Suaedion). Holarctic.
Characteristic species: Salicornia europaea, Suaeda macrocarpa, Suaeda maritima.

25a. Thero-Suaedetalia Br.-Bl. & O. Bolòs 1958
[Thero-Salicornietalia Pignatti 1953 (art. 36) nom. ambig. propos.]
Characteristic species: See alliance (25.1).

25.1. Thero-Suaedion Br.-Bl. in Br.-Bl., Roussine & Nègre 1952
[Thero-Salicornion Br.-Bl. 1933 (art. 36) nom. ambig. propos.]
Characteristic species: Salsola soda, Suaeda albescens, Suaeda spicata, Suaeda splendens.

25b. Thero-Salicornietalia Tüxen in Tüxen & Oberdorfer ex Géhu & Géhu-Franck 1984
Characteristic species: Salicornia emerici, Salicornia obscura.

25.2. Salicornion dolichostachyo-fragilis Géhu & Rivas-Martínez ex Géhu & Géhu-Franck 1984
Characteristic species: Salicornia dolichostachya, Salicornia lutescens.

25.3. Salicornion europaeo-ramosissimae Géhu & Géhu-Franck 1984
Characteristic species: Salicornia pusilla, Salicornia ramosissima.

25.4. Salicornion patulae Géhu & Géhu-Franck 1984
Characteristic species: Salicornia patula.

25.5. Microcnemion coralloidis Rivas-Martínez 1984
Characteristic species: Halopeplis amplexicaulis, Microcnemum coralloides.