Vascular plant communities of Spain and Portugal. Addenda to the Syntaxonomical checklist of 2001

Salvador Rivas-Martínez, Tomás E. Díaz, Federico Fernández-González, Jesús Izco, Javier Loidi, Mario Lousã & Ángel Penas

Itinera Geobotanica 15(1-2): 5-922 (2002)


Go to: Contents - Previous page - Next page

Syntaxon descriptions: Medicagini littoralis - Petrocoptido pyrenaicae

Medicagini littoralis-Stipetum capensis ass. nova hoc loco (39.13.9)

(Taeniathero-Aegilopion geniculatae, Thero-Brometalia, Stellarietea mediae)

Typus associatio: Table 64, rel. 6 [Valencia: Burjassot, YJ2176. 50 m, N, 20 m2].

Characteristic species (territorials): Aegilops geniculata, Medicago littoralis, Rostraria cristata, Stipa capensis.

Diagnosis: Therophytic subnitrophilous grassland dominated by Stipa capensis, Bromus rubens and Aegilops geniculata, in which other thermophilous plants such as Rostraria cristata, Lamarckia aurea and Medicago littoralis are also remarkable. It blooms in the very early spring and grows in waste, barren places of coastal areas in Valencian-Tarraconensean and Setabensean Sectors (Catalan-Valencian Subprovince), mostly in thermomediterranean dry-subhumid bioclimatic belts.

Table 64

39.13.9 Medicagini littoralis-Stipetum capensis
(Taeniathero-Aegilopion geniculatae, Thero-Brometalia, Stellarietea mediae)

Altitude (1=10m) 10 20 20 8 1 5 10 1 7 3 8
Number of species 22 27 25 21 20 31 29 16 23 20 23
Ordinal number 1 2 3 4 5 6* 7 8 9 10 11
Characteristic species:
Stipa capensis 1 3 3 2 1 3 3 1 3 2 V
Bromus rubens 4 3 3 4 3 3 3 2 5 4 V
Rostraria cristata 1 2 + + 1 1 1 + 1 + V
Medicago littoralis + 1 1 1 + + 1 . 1 2 V
Aegilops geniculata 2 1 . 1 + + . 2 1 + IV
Hedypnois cretica + + 1 1 + + + . . + IV
Plantago lagopus + + 1 . 1 1 + 1 + . IV
Lamarckia aurea . 1 + . + + 1 + + + IV
Filago pyramidata + + . 1 . + + . 1 . IV
Lolium rigidum . + + . + + 1 . + . IV
Avena barbata . . + . 1 + + 1 . + IV
Herniaria cinerea + + . + . + + . . . III
Anagallis arvensis . + + . + . + . + . III
Paronychia capitata . + + . . + + . + . III
Papaver rhoeas . + 1 . . + + . . + III
Sonchus tenerrimus . + . . + + . + + . III
Bromus madritensis . . 1 . . 1 . 1 + + III
Companion species:
Medicago minima + . + 1 . 1 + . . + IV
Linum strictum + . + . . + + . + + IV
Leontodon longirostris + . . . . + + + . + III
Limonium echioides + . . + . . . + . + III
Convolvulus arvensis . . . + + . . + . + III

Other species. Characteristic species: Desmazeria rigida + in 1, 2 and 4, 1 in 7. Silene nocturna + in 1 and 9, 1 in 3 and 5. Aegilops triuncialis 1 in 1, + in 3, 7 and 9. Reichardia intermedia + in 1, 4, 7 and 9. Misopates orontium + in 2, 3, 6 and 7. Euphorbia pinea + in 2, 4, 5 and 8. Lobularia maritima 1 in 2, + in 4, 7 and 8. Calendula arvensis + in 3, 4 and 10, 1 in 9. Reseda phyteuma + in 3, 5, 7 and 9. Hordeum leporinum + in 5, 6, 7 and 10. Centaurea melitensis + in 1, 6 and 9. Asphodelus fistulosus + in 2, 3 and 9. Urospermum picroides + in 2 and 7, 1 in 10. Astragalus sesameus + in 1 and 3. Vulpia ciliata + in 1 and 6. Sherardia arvensis + in 3 and 6. Salvia verbenaca + in 3 and 7. Schismus barbatus + in 2. Lagurus ovatus + in 5. Trigonella monspeliaca + in 6. Companion species: Silene sclerocarpa + in 1, 7 and 10. Eryngium campestre + in 2, 3 and 6. Piptatherum miliaceum + in 4, 6 and 8. Phagnalon rupestre + in 1 and 7. Stipa parviflora 1 in 2, + in 7. Plantago albicans + in 4 and 6. Medicago sativa + in 6 and 8. Arenaria leptoclados +, Ononis sicula +, Polycarpon tetraphyllum + in 2. Delphinium gracile +, Hyparrhenia hirta +, Scabiosa stellata + in 4. Dactylis hispanica +, Scabiosa atropurpurea + in 5. Linaria simplex +, Marrubium alysson + in 6. Ononis pubescens +, Silene decipiens + in 9. Melilotus parviflorus + in 10.

Localities: 1. Valencia: Bétera, YJ1988. 29S QE 41. W, 10 m2. 2. Valencia: Llíria, Collado de los Perros, YJ0590. S, 30 m2. 3. Valencia: Bétera, Pla de la Torre, YJ1393. S, 20 m2. 4. Valencia: Massamagrell, El Bogalar, YJ2785. 40 m2. 5. Valencia: Valencia, El Saler, YJ3159. 30 m2. 6. Holotypus ass. Valencia: Burjassot, YJ2176. 50 m, N, 20 m2. 7. Valencia: Sagunto, YJ3395. SE, 30 m2. 8. Valencia: Puçol, motorway A-7 borders, YJ3289. 25 m2. 9. Castellón: Almenara, YK3804. 40 m2. 10. Castellón: Oropesa del Mar, BE5542. E, 50 m2. 11. Synthesized table.

[M.B. Crespo]

Mnio horni-Vandenboschietum speciosae ass. nova hoc loco (30.3.4)

(Hymenophyllion tunbrigensis, Anomodonto-Polypodietalia, Anomodonto-Polypodietea)

Typus associatio: Table 65, rel. 2. [Asturias: Llanes, Valle del río de Nueva. 30TUP385095. NE, 290 m, 1 m2, in Hyperico androsaemi-Alnetum fagetosum sylvaticae].

Characteristic species: Vandenboschia speciosa, Mnium hornum, Diplophyllum albicans, Pseudotaxiphyllum elegans, Jubula hutchinsiae, Lejeunea lamacerina, Lejeunea ulicina.

Diagnosis: Bryo-pteridophytic community that inhabits cavities and small caves on silicic large rock, walls and earthy slopes over a thin layer of soil (exocomophytics) developed in a really wet environement, under the canopy of riverside forests in Southern Cantabrian-Atlantic territories (Alnion incanae: Hyperico androsaemi-Alnetum glutinosae, Valeriano pyrenaicae-Alnetum glutinosae, etc.) in thermo and mesotemperate humid and hyperhumid euoceanic bioclimatic belts. Topographically it gets in contact with Dryopterido aemulae-Hymenophylletum tunbrigensis from which it differs by its floristic composition and its particular habitat: cavities and small caves with thick humic layer.

Table 65

30.3.4 Mnio horni-Vandenboschietum speciosae
(Hymenophyllion tunbrigensis, Anomodonto-Polypodietalia, Anomodonto-Polypodietea)

Altitude (1=10m) 14 29 30 31 31 31 30 32 32 18 18 19 19 19 25
Number of species 6 7 7 9 10 10 5 5 15 11 7 7 7 11 8
Ordinal number 1 2* 3 4 5 6 7 8 9 10 11 12 13 14 15
Characteristic species (territorials):
Vandenboschia speciosa 4 4 3 3 4 5 4 4 4 4 4 4 3 4 V
Mnium hornum 2 1 1 1 1 + 2 . 1 + + 2 1 2 V
Diplophyllum albicans + 1 1 1 2 2 + + 1 + . . . . IV
Pseudotaxiphyllum elegans . 1 2 1 + . + . 1 + + . . + III
Saccogyna viticulosa 1 1 . . 4 . . . 1 + . 2 3 . III
Jubula hutchinsiae . . . + . . . . . . 2 1 2 + II
Lejeunea lamacerina . . . . . 3 . + . 2 . . . + II
Isothecium myosuroides 2 . . . . . . . . 1 . . . 1 II
Plagiochila exigua . . . . . . . + . + . . . . I
Leucobryum juniperoideum . + . . + . . . + . . . . . I
Hymenophyllum tunbrigense . . . . + . . . + . . . . . I
Lejeunea ulicina 1 . . . . 2 . . . . . . . . I
Companion species:
Pellia epiphylla . . 1 1 + + 2 . . . . 2 . . II
Fissidens serrulatus . . . . . . . . 2 . 2 3 3 . II
Oxalis acetosella . . + 1 . . . . 1 + . . . . II
Heterocladium heteropterum . . . . 3 . . . + . 1 . . . I
Saxifraga hirsuta . . 1 2 . . . . + . . . . . I
Hedera helix . . . + . . . . + + . . . . I
Eurhynchium stokesii . . . . . . . . + . . . . + I
Conocephalum conicum . . . . . . . . . . . + . 2 I
Dumortiera hirsuta . . . . . . . . . . 1 . 3 . I
Hylocomium armoricum . . . . + . . + . . . . . . I

Other species. Characteristic species: Bazzania trilobata + in 2. Calypogeia muelleriana + in 6. Companion species: Dryopteris aemula + in 4. Fissidens polyphyllus 1, Heterocladium fallax 1, Blechnum spicant + in 6. Fissidens taxifolius + in 8. Woodwardia radicans 1, Plagiochila porelloides + in 9. Plagiothecium nemorale + in 10. Mnium marginatum + in 13. Hookeria lucens 3, Chiloscyphus fragans 1, Eurhynchium striatum 1 in 14.

Localities: 1. Asturias, Llanes: Valle río de Nueva. 30TUP405095. NW, 1 m2 in Hyperico androsaemi-Alnetum osmundetosum regalis. 2. Holotypus ass. Asturias, Llanes: Valle río de Nueva. 30TUP385095. NE, 1 m2 in Hyperico androsaemi-Alnetum fagetosum sylvaticae. 3, 4. Asturias, Llanes: Valle río de Nueva. 30TUP385090. NE-W, 0,9 m2 in Hyperico androsaemi-Alnetum fagetosum sylvaticae. 5, 6, 7. Asturias, Llanes: Valle río de Nueva. 30TUP380090. N-NE, 0,5 m2 in Hyperico androsaemi-Alnetum fagetosum sylvaticae. 8, 9. Asturias, Llanes: Valle río de Nueva. 30TUP375090. SW-W, 0,5 m2 in Hyperico androsaemi-Alnetum fagetosum sylvaticae. 10, 11, 12, 13, 14. Asturias, Llanes: Valle río de Nueva. 30TUP405095. N, 0,5 m2 in Hyperico androsaemi-Alnetum osmundetosum regalis. 15. Synthesized table.

[T.E. Díaz, M.C. Fernández & Collado]

Myriophyllo alterniflori-Potametum natantis ass. nova hoc loco (3.2.1)

(Nymphaeion albae, Potametalia, Potametea)

Typus associatio: Table 66, rel. 5 [Avila: Navalsauz, Puente Nuevo, Alberche river. 30TUK2772. 1240 m, 2 m2].

Characteristic species (territorials): Myriophyllum alterniflorum, Potamogeton natans, Ranunculus pseudofluitans.

Diagnosis: Aquatic carpets of nymphaeids (Potamogeton natans) and myriophyllids, often accompanied by batrachids or small elodeids, growing in slow running or standing, moderately deep and mesotrophic water bodies (50-200 cm maximum depth) associated to river courses or ponds subjected to strong fluctuations of the water level but that rarely undergo a complete dessication. The association is known from the rivers of the north and south slopes of the Iberian Central System, but it has probably a wider meso-supramediterranean, Mediterranean West Iberian distribution.

Table 66

3.2.1 Myriophyllo alterniflori-Potametum natantis
(Nymphaeion albae, Potametalia, Potametea)

Altitude (1=10m) 124 122 122 124 124 122 122 108 12
Number of species 3 3 3 4 4 4 4 7 4
Ordinal number 1 2 3 4 5* 6 7 8 9
Characteristic species:
Potamogeton natans 3 5 5 5 5 4 4 4 V
Myriophyllum alterniflorum 2 2 3 2 3 3 1 2 V
Ranunculus pseudofluitans . + 1 + + 1 1 . IV
Potamogeton berchtoldii . . . 1 2 2 . . III
Companion species:
Alopecurus aequalis 3 . . . . . 1 . II

Other species. Companion species: Lemna minor 1, Utricularia minor 1, Callitriche brutia +, Rorippa nasturtium-aquaticum +, Sparganium microcarpum + in 8.

Localities: 1. Ávila: San Martín del Pimpollar, Alberche river from Venta del Obispo to Piquillo river. 30TUK2873. 4 m2. 2. Ávila: San Martín del Pimpollar, Alberche river from Venta del Obispo to Piquillo river. 30TUK2872. 2 m2. 3. Ávila: San Martín del Pimpollar, Alberche river from Venta del Obispo to Piquillo river. 30TUK2872. 2 m2. 4. Ávila: Navalsauz, río Alberche, Puente Nuevo. 30TUK2774. 2 m2. 5. Holotypus ass. Ávila: Navalsauz, Alberche river, Puente Nuevo. 30TUK2772. 2 m2. 6. Ávila: Navalsauz, Puente Nuevo, Alberche river. 30TUK2772. 2 m2. 7. Ávila: San Martín del Pimpollar, Alberche river from Venta del Obispo to Piquillo river. 30TUK2873. 4 m2. 8. Ávila: Umbrías, Aravalle, Retuerta dam. 30TTK8067. Deep = 200 cm, 10 m2. 9. Synthesized table.

[Rivas-Martínez, Fernández-González, Sánchez-Mata, Pizarro & Sardinero]

Onopordenea acanthii subclassis nova hoc loco (34B)

[Onopordenea acanthii Rivas-Martínez, Báscones, T.E. Díaz, Fernández-González & Loidi in Itinera Geobot. 5: 374. 1991 (art. 8, 17), Onopordetea Br.-Bl., Pflanzensoziologie: 131. 1964 (corresp. name)]
(Artemisietea vulgaris)

Typus subclassis: Onopordetalia acanthii Br.-Bl. & Tüxen ex Klika & Hadač 1944.

Noticeable characteristic species: Artemisia absinthium, Carduus nutans, Carduus pycnocephalus, Carduus tenuiflorus, Centaurea calcitrapa, Centaurea solstitialis, Chondrilla juncea, Echium vulgare, Eryngium campestre, Hyoscyamus niger, Lactuca virosa, Onopordum acanthium, Picnomon acarna, Reseda luteola, Verbascum pulverulentum, Verbascum thapsus, Verbascum virgatum.

Diagnosis: Nitrophilous and subnitrophilous vegetation from turned over soils or animal resting places, where the great thistles are preponderant, as well as other annual, biennal or rosulate perennial plants. It flowers in late spring or summer and has a Mediterranean and Eurosiberian optimum, appearing in a depauperate form, sometimes as a neophytic, in other Holarctic or tropical regions with a somewhat warm and dry summer (Saharian-Arabian, Punan, etc.). To the moment, two orders are recognized in the subclassis: Onopordetalia acanthii, nomenclatural type of the subclassis, with an Eurosiberian areal and preference for the continental or submediterranean climates, relatively poor in characteristic species and floristicaly and successionaly rather related to the nitrophilous perennial vegetation of the Arction (Artemisietalia vulgaris), and the order Carthametalia lanati, with Mediterranean optimum, rich in characteristic species and from our point of view, sinecological type and dispersal center of the subclassis.

[Rivas-Martínez, Báscones, T.E. Díaz, Fernández-González & Loidi]

Onopordetum acantho-castellani ass. nova hoc loco (34.10.10)

(Onopordion castellani, Carthametalia lanati, Onopordenea acanthii, Artemisietea vulgaris)

Typus associatio: Table 67, rel. 10 [Valladolid: Villardefrades. 30TUM12. 20 m2].

Characteristic species (territorials): Onopordum acanthium, Onopordum nervosum subsp. castellanum, Onopordum xglomeratum (O. acanthium x O. nervosum subsp. castellanum).

Diagnosis: Castilian meso-supramediterranean dry clayey calcareous subnitrophilous frequent community well developed on marly soils in disturbed areas by human impacts, such as spoil banks, field crop environs, orchards, off-road places, and other sites near the villages. This community can be differentiated of Aragonese and Sorian-Oroiberian association Onopordetum castellani by Carduus granatensis, Echium salmanticum and Verbascum pulverulentum and, in opposite sense by Carduus assoi, Carduus nigrescens and Onopordum corymbosum.

Table 67

34.10.10 Onopordetum acantho-castellani
(Onopordion castellani, Carthametalia lanati, Onopordenea acanthii, Artemisietea vulgaris)

Altitude (1=10m) 69 78 77 77 108 84 90 73 83 78 71 91 73 76 84 81 81
Number of species 12 13 13 14 15 15 16 16 16 16 16 17 17 17 19 19 16
Ordinal number 1 2 3 4 5 6 7 8 9 10* 11 12 12 14 15 16 17
Characteristic species:
Onopordum castellanum 3 2 3 3 3 3 3 3 3 4 4 2 3 3 4 3 V
Cichorium intybus 2 1 + 2 2 2 2 1 1 1 1 2 + 2 1 + V
Onopordum acanthium 1 1 1 + . + 2 1 + + 1 + 1 + + + V
Scolymus hispanicus 1 2 1 . 2 . 2 1 2 + + 2 2 1 + 1 V
Carthamus lanatus 1 3 1 1 2 . 2 2 + + . + 1 1 + 2 V
Carduus tenuiflorus 1 1 1 1 . + . 1 + + + . + 1 . + IV
Lactuca serriola + . . 1 1 + . . . 2 1 + + 1 1 1 IV
Verbascum pulverulentum + . + + . + 1 1 + . . 1 . + . 2 IV
Chondrilla juncea . 1 . . . + 1 1 1 . + . . . 1 + III
Eryngium campestre . . . . 2 . 1 . . . . 1 1 . 1 1 II
Echium asperrimum . . . + . . . 1 1 . + . + + . . II
Verbascum montanum 1 . 1 . . . . . . . + . . + + + II
Melilotus altissimus + . . . . . . . . . + . + + . . II
Picnomon acarna . + 2 . 1 . . . . 1 . . . . . . II
Carduus granatensis . . . 1 . 2 1 . . . . . . . 1 . II
Echium vulgare . . . + . . . 1 2 . . . . . . . I
Echium salmanticum . . . 1 . 1 . . . . . . . . + . I
Companion species:
Hordeum leporinum 1 2 2 1 2 1 1 2 + + 1 2 1 1 1 2 V
Avena sterilis . 2 + . . 1 1 1 1 1 + 1 1 1 1 1 V
Hirschfeldia incana 1 2 . . 1 . 1 2 2 1 + + . + + 2 IV
Bromus diandrus 1 1 1 . . + . . . 1 . . . + . 1 III
Cirsium arvense . . . 1 2 . . 1 . + 1 2 . 1 . + III
Cirsium pyrenaicum . . + 1 + + 1 . . . . 2 . . + 1 III
Phlomis herba-venti . . . . 2 1 . 1 1 . . 1 . . . . II
Anacyclus clavatus . . 1 . . . 1 . . . . 1 2 + . 1 II

Other species. Characteristic species: Carduus bourgaeanus 2 in 3, 1 in 14. Foeniculum piperitum 2 in 8, 1 in 13. Onopordum xglomeratum + in 10 and 11. Onopordum xcostae + in 13 and 15. Onopordum illyricum 2 in 2. Carlina hispanica 1, Centaurea calcitrapa 1 in 7. Silybum marianum + in 11. Lactuca virosa 1 in 12. Verbascum virgatum 1 in 14. Cirsium odontolepis 1 in 15. Cirsium vulgare + in 16. Companion species: Daucus carota 2 in 5, + in 12. Mantisalca salmantica 1 in 13, + in 15. Medicago sativa 1 in 13, + in 16. Dipsacus sylvestris 1 in 5. Anchusa italica 1 in 9. Euphorbia serrata + in 11. Brachypodium phoenicoides 1 in 15.

Localities: 1. Zamora: Cerecinos de Campos. 30TTM44. 20 m2. 2. Valladolid: Medina del Campo. 30TUL37. 20 m2. 3. Palencia: Palencia. 30TUM75. 40 m2. 4. Palencia: Torremormojón. 30TUM55. 20 m2. 5. Segovia: Ayllón. 30TVL68. 40 m2. 6. Valladolid: Villanubla. 30TUM42. 20 m2. 7. Valladolid: Cogeces del Monte. 30TUL89. 70 m2. 8. Valladolid: Valladolid. 30TUM51. 20 m2. 9. Ávila: Arévalo. 30TUL54. 20 m2. 10. Holotypus ass. Valladolid: Villardefrades. 30TUM12. 20 m2. 11. Valladolid: Tordesillas. 30TUL39. 20 m2. 12. Soria: Aldea de S. Esteban. 30TVM70. 40 m2. 13. Valladolid: Simancas. 30TUM40. 40 m2. 14. Valladolid: Mota del Marqués. 30TUM11. 40 m2. 15. Valladolid: Torrelobatón. 30TUM21. 40 m2. 16. Burgos: Torregalindo. 30TVL38. 40 m2. 17. Synthesized table.

[Rivas-Martínez & Sánchez-Mata]

Orlayo grandiflorae-Aegilopetum geniculatae Romo ass. nova hoc loco (39.13.11)

[Aegilopo-Orlayetum grandiflorae Romo in Arxius Secc. Ci. Inst. Estud. Catalans 90: 408, tb. 16. 1989 (art. 5)]
(Taeniathero-Aegilopion geniculatae, Thero-Brometalia, Stellarietea mediae)

Typus associatio: Aegilopo-Orlayetum grandiflorae Romo in Arxius Secc. Ci. Inst. Estud. Catalans 90: 408, tb. 16, rel. 2. 1989. Lleida: Puente de Montañana. CG0969. 535 m, 50 m2. (Holotypus hoc loco, l.c. rel. 2). Characteristic species: 4 Aegilops geniculata, 1 Orlaya grandiflora, + Bromus hordeaceus, + Medicago orbicularis, + Petrorhagia prolifera. Companion species: + Alyssum alyssoides, + Hypericum perforatum, + Phleum phleoides, + Phlomis herba-venti, + Sanguisorba muricata.

Characteristic species (territorials): Aegilops geniculata, Orlaya grandiflora.

Diagnosis: Prepyrenean annual ephemerous early summer blooming calcicolous grassland-like community spread in upper mesotemperate and lower supratemperate submediterranean subhumid altered zones.

[Rivas-Martínez & Izco]

Paeonio broteroi-Abietion pinsapo (Rivas-Martínez 1987) all. nova, stat. nov. hoc loco (76.11)

[Paeonio-Abietenion pinsapo Rivas-Martínez, Mem. Mapa Series Veg. España: 160. 1987 (art. 27a), Paeonio-Abietion pinsapo Rivas-Martínez in Ecol. Medit. 8: 284. 1982 (art. 8)]
(Quercetalia humilis, Querco-Fagetea)

Typus alliancia: Paeonio broteroi-Abietetum pinsapo Asensi & Rivas-Martínez in Anales Inst. Bot. Cavanilles 33: 243, 1976 (art. 27a). Typus associatio (holotypus): Asensi & Rivas-Martínez, l.c., tb. 1, rel. 1.

Characteristic species: Abies pinsapo, Bunium macuca, Ononis reuteri.

Diagnosis: Supra-mesomediterranean humid to hyperhumid pinsapo fir forests (Abies pinsapo), growing in soils developed on calcitic dolomite, or ultramafic igneous rocks in Rondean Biogeographic Sector (Betic Province). Bioclimatic diagnosis: Ic 15-17, I0 9.0-18.0, It 100-230, Tps 500-600.

Bunio macucae-Abietetum pinsapo (Asensi & Rivas-Martínez 1976) Rivas-Martínez, Mem. Mapa Series Veg. España: 160. 1987 (76.11.1)

Diagnosis: Ultramafic Bermejan Rondean pinsapo fir (Abies pinsapo) forest community.

Paeonio broteroi-Abietetum pinsapo Asensi & Rivas-Martínez in Anales Inst. Bot. Cavanilles 33: 243, tb. 1. 1976 (76.11.2)

Diagnosis: Dolomite and calcitic dolomite Rondean pinsapo fir (Abies pinsapo) forest community.

[Rivas-Martínez & Asensi]

Parafestucetalia albidae ordo novus hoc loco (57b)

[Festucetalia jubatae Rivas-Martínez in Capelo, J.C. Costa, Lousã, Fontinha, Jardim, Sequeira & Rivas-Martínez in Silva Lusit. 7(2): 275. 2000 (art. 3b)]
(Stipo giganteae-Agrostietea castellanae)

Typus ordo: Deschampsio maderensis-Parafestucion albidae Capelo, J.C. Costa, Lousã, Fontinha, Jardim, Sequeira & Rivas-Martínez in Silva Lusit. 7(2): 275. 2000.

Characteristic species: Agrostis obtusissima, Anthyllis lemanniana, Armeria maderensis, Crepis vesicaria subsp. andryaloides, Parafestuca albida.

Diagnosis: Madeiran silicicolous perennial grasslands growing on shallow soils and rupestrian biotopes in meso-supratemperate hyperhumid and ultrahyperhumid summit areas of Madeira Isle in contact with the climactical zone of Erica arborea microforest (Polysticho falcinelli-Ericetum arboreae).

[Rivas-Martínez, Capelo, J.C. Costa, Lousã, Fontinha, Jardim & Sequeira]

Parietarion lusitanico-mauritanicae all. nova hoc loco (41.3)

Typus alliancia: Torilido nodosae-Parietarietum mauritanicae Rivas-Martínez in Anales Inst. Bot. Cavanilles 34(2): 564, tb. 4, rel. 1 (holotypus). 1978.

Characteristic species: Ceratocapnos heterocarpa, Galium minutulum, Mercurialis elliptica, Parietaria lusitanica, Parietaria mauritanica, Stachys circinata, Succowia balearica, Theligonum cynocrambe, Urtica dubia.

Diagnosis: Annual spring ephemeral internal and external wood, shrub fringes and walls slightly nitrified semi-shaded communities spread in thermo- and lower mesomediterranean semiarid to subhumid in West Mediterranean.

Anogrammo leptophyllae-Parietarietum lusitanicae Rivas-Martínez & Ladero in Rivas-Martínez in Anales Inst. Bot. Cavanilles 34(2): 562, tb. 3. 1978 (41.3.1)

Distribution: Lusitan-Extremadurean lower mesomediterranean dry to subhumid growing on wooded and shaded granit caos.

Castellio tuberculosae-Geranietum rotundifolii Alcaraz, Garre, Martínez-Parras & Peinado in Collect. Bot. (Barcelona) 16(2): 417, tb. 2. 1986 (41.3.2)

Distribution: Almeriensian coastal semi-shaded community under shrubs of Rhamno-Juniperetum turbinatae.

Fumario macrosepalae-Parietarietum mauritanicae J.M. Losa & Pérez-Raya in J.M. Losa in Lazaroa 10: 24, tb. 1. 1988 (41.3.3)

Distribution: Lower mesomediterranean Granatensean Celtis australis riparian and gorges forests.

Geranio purpurei-Galietum minutuli Rivas-Martínez, Costa, Castroviejo & E. Valdés in Lazaroa 2: 74, tb. 39. 1980 (41.3.4)

Distribution: Gaditan-Coastal Onubensean shaded community growing on tangel humus of Osyrio-Juniperetum turbinatae littoral dune bush.

Geranio rotundifolii-Silenetum latifoliae O. Bolòs, Folch & Vigo in O. Bolòs in Folia Bot. Misc. 6: 126, tb. 6. 1989 (41.3.5)

Distribution: Known from Columbretes Isles on shady rocks.

Geranio rotundifolii-Theligonetum cynocrambes Rivas-Martínez & Malato-Beliz in Rivas-Martínez 1978 corr. hoc loco (41.3.6)

[Geranio pusilli-Theligonetum cynocrambes Rivas-Martínez & Malato-Beliz in Rivas-Martínez in Anales Inst. Bot. Cavanilles 34(2): 568, tb. 5, holotypus rel. 5. 1978 (art. 43)]

Distribution: Thermomediterranean calcicole growing in the shade of Rhamno oleoidis-Quercetum rotundifoliae trees.

Mercuriali ambiguae-Succowietum balearicae O. Bolòs, Folch & Vigo in O. Bolòs in Folia Bot. Misc. 6: 125, tb. 5. 1989 (41.3.7)

Distribution: Known from Columbretes Isles in the shade of walls.

Mercuriali ellipticae-Theligonetum cynocrambes Peinado, Martínez-Parras & Bartolomé in Studia Bot. 5: 64, tb. 5. 1986 (41.3.8)

Distribution: Thermomediterranean calcicolous Rondean in the shade of walls.

Parietario mauritanicae-Ceratocapnetum heterocarpae Martínez-Parras in Anales Jard. Bot. Madrid 39(1): 187, tb. 1. 1982 (41.3.10)

Distribution: Betic and Almeriensian thermomediterranean dry and semiarid, characterized by Ceratocapnos heterocarpa, growing in the shade of the trees and shrubs of Quercetea ilicis communities.

Soncho dianae-Parietarietum lusitanicae Esteve , Veget. Fl. prov. Murcia: 85. 1973 (41.3.11)

[Parietario lusitanicae-Geranietum purpurei Alcaraz, Garre, Martínez-Parras & Peinado in Collect. Bot. (Barcelona) 16(2): 416, tb. 1. 1986, accepted in Itinera Geobot. 14: 97. 2001 (41.3.9) (syntax. syn.)]

Distribution: Thermo-lower mesomediterranean semiarid Murcian-Almeriensian growing in the shade of shrubs of Pistacio-Rhamnetalia alaterni communities.

Theligono cynocrambes-Veronicetum cymbalariae O. Bolòs, Molinier & P. Montserrat in Acta Geobot. Barcinon. 5: 97. 1970 (41.3.12)

Distribution: Minorcan in the shade of walls.

Torilido nodosae-Parietarietum mauritanicae Rivas-Martínez in Anales Inst. Bot. Cavanilles 34(2): 564, tb. 4. 1978 (41.3.13)

Distribution: Lusitan-Extremadurean lower mesomediterranean dry and subhumid growing in the shade of trees and shrubs of Quercion broteroi and Asparago-Rhamnion communities.

[Rivas-Martínez & Cantó]

Petrocoptido pyrenaicae-Sarcocapnetea enneaphyllae classis nova hoc loco (29)

Typus classis (holotypus): Petrocoptidetalia pyrenaicae ordo novus hoc loco.

Characteristic species: Asplenium csikii, Sarcocapnos enneaphylla.

Diagnosis: Overhanging rock crevice chasmophytic ombrophobic communities which grow on limestone or dolomite rich rocks in Mediterranean and Temperate mountain cliffs and canyons of Spanish Iberian Peninsula and North African Mauritania.

Sarcocapnetalia enneaphyllae F. Casas in Trab. Dep. Bot. Univ. Granada 1: 22. 1972 (29a)

Typus ordo (lectotypus: art. 20): Sarcocapnion enneaphyllae F. Casas in Trab. Dep. Bot. Univ. Granada 1: 30. 1972.

Characteristic species: Asplenium celtibericum subsp. celtibericum.

Diagnosis: Overhanging rock crevice chasmophyte ombrophobic communities of Spanish Mediterranean Iberian Peninsula and North African Mauritania in thermo to supramediterranean pluviseasonal and xeric bioclimates and occasionaly temperate submediterranean.

Sarcocapnion enneaphyllae F. Casas in Trab. Dep. Bot. Univ. Granada 1: 30. 1972 (29.1)

Typus alliancia (lectotypus hoc loco): Antirrhinetum pertegasii O. Bolòs in Mem. Real Acad. Ci. Barcelona 38(1): 9. 1967, excl. typus: Sarcocapnetum enneaphyllae Rivas Goday 1941 (art. 2b, 7) ap. F. Casas in Trab. Dep. Bot. Univ. Granada 1: 30. 1972.

Characteristic species: Antirrhinum pertegasii (29.1.2), Antirrhinum pulverulentum (29.1.3), Antirrhinum microphyllum (29.1.1), Chaenorhinum origanifolium subsp. semiglabrum (29.1.7), Chaenorhinum segoviense (29.1.6), Moehringia castellana (29.1.8).

Diagnosis: Meso-supramediterranean Catalan-Valencian and Mediterranean Central Iberian calcareous and dolomitic limestones overhanged and walls rather nitrified communities.

Antirrhinetum microphylli F. Casas in Cuad. Ci. Biol. (Granada) 3: 92, tb. 3. 1974 (29.1.1)

Distribution: Alcarrean upper mesomediterranean dry on calcareous and dolomitic overhangs, characterized by Antirrhinum microphyllum.

Antirrhinetum pertegasii O. Bolòs in Mem. Real Acad. Ci. Barcelona 38(1): 9, tb. 2. 1967 (29.1.2)

Distribution: Tarraconensean supramediterranean subhumid on calcareous overhangs, characterized by Antirrhinum pertegasii.

Antirrhinetum pulverulenti F. Casas in Cuad. Ci. Biol. (Granada) 3: 91, tb. 2. 1974 (29.1.3)

Distribution: Celtiberian-Alcarrean and East Maestracensean supramediterranean dry and subhumid on calcareous overhangs, characterized by Antirrhinum pulverulentum.

Asplenio pachyrachidis-Sarcocapnetum enneaphyllae F.J. Pérez, T.E. Díaz & P. Fernández in Monogr. Inst. Piren. Ecol. (Jaca) 5: 571, tb. 1. 1990 (29.1.4)

Distribution: Castilian Cantabrian supramediterranean subhumid on calcareous overhangs, diferenciated by Chaenorhinum origanifolium subsp. origanifolium.

Chaenorhino crassifolii-Sarcocapnetum enneaphyllae Rivas-Martínez & G. López in G. López in Anales Inst. Bot. Cavanilles 34(2): 611, tb. 5. 1978 (29.1.5)

Distribution: Alcarrean and Manchean meso-supramediterranean dry and subhumid on calcareous nitrified overhangs, characterized by Chaenorhinum origanifolium subsp. crassifolium.

Chaenorhino segoviensis-Sarcocapnetum enneaphyllae Loidi & Galán in Studia Bot. 7: 163, tb. 3. 1989 (29.1.6)

Distribution: Celtiberian-Alcarrean upper meso-supramediterranean dry on calcareous nitrified overhang walls, characterized by Chaenorhinum segoviense subsp. segoviense.

Chaenorhino semiglabri-Asplenietum celtiberici G. Navarro in Opusc. Bot. Pharm. Complut. 5: 46, tb. 20. 1989 (29.1.7)

Distribution: Sorian Oroiberian supratemperate submediterranean humid on calcareous overhangs, characterized by Chaenorhinum segoviense subsp. semiglabrum.

Moehringietum castellanae ass. nova hoc loco (29.1.8)

Typus associatio: Table 68, rel. 1 [Guadalajara: Alpedrete de la Sierra, Pontón de la Oliva, Quebrantahuesos area, overhang limestone wall. 40º 51'N-3º 27'W. 850 m, NW, 20 m2].

Characteristic species: Moehringia castellana.

Diagnosis: Celtiberian-Alcarrean lower and East Maestracensean, supramediterranean dry and subhumid on calcareous north exposed overhangs (Serranía de Cuenca, Duratón river gorges and Patones-Pontón de la Oliva cliffs), characterized by Moehringia castallana.

Table 68

29.1.8 Moehringietum castellanae
(Sarcocapnion enneaphyllae, Sarcocapnetalia, Petrocoptido-Sarcocapnetea)

Altitude (1=10m) 85 86 86 85 85 85
Number of species 5 7 6 6 7 6
Ordinal number 1* 2 3 4 5 6
Characteristic species:
Moehringia castellana 2 2 2 3 2 V
Sarcocapnos enneaphylla 1 3 2 1 2 V
Chaenorhinum segoviense . . . + . I
Asplenium csikii . . . . + I
Companion species:
Rhamnus pumilus + + + 1 1 V
Chiliadenus saxatilis + + 1 . . III
Parietaria judaica + + . . + III
Sedum dasyphyllum . . + + + III

Other species. Companion species: Ficus carica +, Umbilicus rupestris + in 2. Sedum album + in 3. Seseli montanum + in 4. Lactuca perennis + in 5.

Localities: 1. Holotypus ass. Guadalajara: Alpedrete de la Sierra, Pontón de la Oliva, Quebrantahuesos area, overhang limestone wall. 40º 51'N-3º 27'W. NW, 20 m2. 2. Guadalajara: Pontón de la Oliva, Maracaibo area. W, 10 m2. 3. Guadalajara: Pontón de la Oliva, Púrpura area. NW, 20 m2. 4. Guadalajara: Pontón de la Oliva, Poyero area. W, 10 m2. 5. Guadalajara: Pontón de la Oliva, Maracaibo area, overhang limestone wall. W, 20 m2. 6. Synthesized table.

Sarcocapnion pulcherrimae F. Casas 1972 corr. Rivas-Martínez, Cantó & Izco hoc loco (29.2)

[Sarcocapnion crassifoliae F. Casas in Trab. Dep. Bot. Univ. Granada 1: 26. 1972 (art. 43)]

Typus alliancia: Sarcocapnetum crassifoliae Cuatrecasas ex Esteve & F. Casas in Cuad. Ci. Biol. 1: 68. 1971.

Characteristic species: Antirrhinum mollissimum (29.2.9), Chaenorhinum tenellum (29.2.6), Gypsophila montserratii (29.2.1), Moehringia giennensis (29.2.3), Moehringia tejedensis (29.3.2), Rupicapnos decipiens (29.2.4),.Sarcocapnos baetica (29.2.8), Sarcocapnos crassifolia subsp. speciosa, Sarcocapnos integrifolia (29.2.5), Sarcocapnos pulcherrima (29.2), Sarcocapnos saetabensis (29.2.6).

Diagnosis: Thermo to supramediterranean on overhanging dolomite, limestone and calcium carbonate rocks, cliffs and canyons of Betic Ranges: Betic and Setabensean biogeographic territories.

Gypsophiletum montserratii F. Casas in Publ. Inst. Biol. Aplicada 52: 123. 1972 (29.2.1)

Distribution: Subbetic Alcaracensean supramediterranean subhumid on calcareous overhangs, characterized by Gypsophila montserratii.

Hieracio texedensis-Moehringietum tejedensis Mota, Gómez-Mercado & F. Valle in Vegetatio 94: 105, tb. 3. 1991 (29.2.2)

Distribution: Malacitan-Texedan upper supramediterranean humid growing on shadow dolomitic overhanged walls, characterized by Hieracium texedense and Moehringia texedensis.

Moehringietum giennensis F. Casas 1972 corr. Mota, Gómez-Mercado & F. Valle in Vegetatio 94: 108. 1991 (29.2.3)

[Moehringietum intricatae F. Casas in Trab. Dep. Bot. Univ. Granada 1: 27, tb. 2. 1972]

Distribution: Subbetic supramediterranean subhumid on dry overhanged calcareous walls, characterized by Moehringia intricata subsp. giennensis.

Rupicapnetum decipientis A.V. Pérez, Cabezudo & Nieto in Acta Bot. Malacitana 20: 311, tb. 1. 1995 (29.2.4)

Distribution: Rondean, Hispalensean and Malacitan thermomediterranean dry to subhumid on calcareous shadow overhanged walls, characterized by Rupicapnos decipiens.

Sarcocapnetum integrifoliae F. Casas & Molero Brion. in F. Casas in Trab. Dep. Bot. Univ. Granada 1: 26, tb. 1. 1972 (29.2.5)

Distribution: Maginan and Sagran Subbetic mesomediterranean dry on overhanged dolomitic limestones, characterized by Sarcocapnos integrifolia.

Sarcocapno saetabensis-Chaenorhinetum tenelli ass. nova hoc loco (29.2.6) (*)

[ass. Chaenorhinum tenellum et Sarcocapnos enneaphylla Rivas Goday, Esteve, Rigual & Borja in Anales Inst. Bot. Cavanilles 12: 490. 1954 (art. 3b), ass. Sarcocapnos crassifolia et Chaenorhinum tenellum (Rivas Goday & Borja 1954) Rigual & Esteve in Rigual, Esteve & Rivas Goday 1963 (art. 7)]

Typus associatio: Valencia: Ayora, pr. Cueva Horadada, 30SXJ8121, 1000 m, 1-VI-1996, 10 m2, cob. 50%, North orientated lime overhang "balma". Characteristic species: 3 Chaenorhinum tenellum, 2 Sarcocapnos saetabensis, 1 Chaenorhinum crassifolium, + Crepis albida (subsp. longicaulis), 1 Hieracium loscosianum, + Chiliadenus saxatilis (= Jasonia saxatilis), + Hypericum ericoides, + Sanguisorba rupicola, + Melica minuta, + Centaurea spachii.

Characteristic species: Chaenorhinum tenellum, Sarcocapnos saetabensis.

Diagnosis: Rupicolous plant community growing on ceilings of calcareous caves and caverns ("balmas"). Chaenorhinum tenellum and Sarcocapnos saetabensis (two remarkable endemics of Eastern Spain) are dominant and other Iberolevantine plants such as Chaenorhinum origanifolium subsp. crassifolium, Hieracium loscosianum, Jasonia saxatilis and Hypericum ericoides are also important. This association is endemic to Setabensean Sector (Catalan-Valencian Subprovince), basically in mesomediterranean subhumid bioclimatic belt.

[M.B. Crespo] (*)

Sarcocapnetum pulcherrimae Cuatrecasas ex Esteve & F. Casas 1971 corr. hoc loco (29.2.7)

[Sarcocapnetum crassifoliae Cuatrecasas ex Esteve & F. Casas in Cuad. Ci. Biol. (Granada) 1: 68. 1971 (art. 43)]

Distribution: Almijarensean, Granatensean and Maginan Subbetic meso-supramediterranean dry and subhumid calcareous and dolomitic-limestones overhangs, characterized by Sarcocapnos pulcherrima.

Sarcocapno baeticae-Centaureetum clementei Asensi & Esteve in Trab. Dep. Bot. Univ. Granada 4: 32, tb. 1977 (29.2.8)

Distribution: Rondean mesomediterranean dry and subhumid calcareous and dolomitic overhangs from Junquera to Grazalema mountains, characterized by Centaurea clementei.

Sarcocapno enneaphyllae-Antirrhinetum mollissimi F. Casas in Publ. Inst. Biol. Aplicada 50: 49, tb. 1. 1971 (29.2.9 = 29.1.9)

Distribution: Almeriensian and Gadorensian thermo-mesomediterranean semiarid and dry on urban walls and overhanged calcareous rocks, characterized by Antirrhinum mollisimum.

Petrocoptidetalia pyrenaicae ordo novus hoc loco (29b)

Typus ordo: Valeriano longiflorae-Petrocoptidion F. Casas in Trab. Dep. Bot. Univ. Granada 1: 28. 1972.

Characteristic species: Petrocoptis sps., Asplenium celtibericum subsp. molinae.

Diagnosis: Pyrenean and Orocantabric overhanging rock crevice chasmophyte ombrophobic communities mostly on temperate and submediterranean North Spain canyons and mountains from meso to orotemperate.

Valeriano longiflorae-Petrocoptidion F. Casas in Trab. Dep. Bot. Univ. Granada 1: 28. 1972 (29.3)

Typus alliancia: Petrocoptidetum crassifoliae O. Bolòs & P. Montserrat ex F. Casas in Ars Pharm. 11: 274. 1970 (art. 43) [Petrocoptidetum hispanicae O. Bolòs & P. Montserrat ex F. Casas 1970 corr. Rivas-Martínez, Cantó & Izco hoc loco].

Characteristic species: Androsace cylindrica (29.3.11), Androsace willkommii (29.3.9), Antirrhinum molle, Antirrhinum sempervirens, Asplenium seelosi subsp. catalaunicum (29.3.2), Petrocoptis crassifolia (29.3.3), Petrocoptis pardoi subsp. guarensis (29.3.14), Petrocoptis pyrenaica subsp. hispanica (29.3.5), Petrocoptis montserratii (29.3.6), Petrocoptis pardoi subsp. montsicciana (29.3.10), Petrocoptis pardoi subsp. pardoi (29.3.8), Petrocoptis pseudoviscosa (29.3.4), Petrocoptis pyrenaica (29.3.7), Valeriana longiflora.

Diagnosis: Meso to orotemperate frequently submediterranean on Prepyrenean canyons, exceptionally on Maestratian Catalanidic South of Ebro.

In some overhangs and caves of Prepyrenean gorges of the Cinca river there are other associations caracterized by their endemics of the genus Petrocoptis A. Braun ex Endl., as Asplenio csikii-Petrocoptidetum crassifoliae [char.: Petrocoptis crassifolia Rouy, Devotas Pass, Cinca river and Cañón de Añisclo], Asplenio csikii-Petrocoptidetum pseudoviscosae [char.: Petrocoptis pseudoviscosa Fern. Casas, Congostos de Ventamillo and de Campo in the Ésera river], Valeriano-Petrocoptidetum guarensis [char.: Petrocoptis guarensis Fern. Casas, Barranco de Mascún and other limestone gorges of the basin of the Alcanadre river in Guara range], Petrocoptido montsiccianae-Antirrhinetum mollis [char.: Petrocoptis montsicciana O. Bolòs & Rivas Mart., in the Prepyrenean gorges of the rivers Noguera Pallaresa and Segre: Congosts of Terradets, Collegats and Oliana].

Antirrhino sempervirentis-Potentilletum alchimilloidis Rivas Goday in Rivas Goday, Esteve, Rigual & Borja in Anales Inst. Bot. Cavanilles 12(1): 487, tb. 4. 1954 (29.3.1)

Distribution: Pyrenean supra-orotemperate subhumid to humid growing on calcareous and calc-schist walls, characterized by Antirrhinum sempervirens. It could be also classified in Saxifragion mediae alliance.

Asplenietum catalaunici F. Casas 1970 corr. O. Bolòs & Vigo, Flora dels Països Catalans 1: 65. 1984 (29.3.2)

[Asplenietum celtiberici F. Casas in Ars. Pharm. 11: 275, tb. 8. 1970 (art. 43)]

Distribution: Eastern Pyrenean High Berguedan supra-oromediterranean humid calcareous overhangs from Cadí-Moixeró Natural Park to Montgrony mountains, characterized by Asplenium seelosi subsp. catalaunicum.

Asplenio csikii-Petrocoptidetum crassifoliae ass. nova hoc loco (29.3.3) (*)

[Petrocoptidetum crassifoliae F. Casas in Cuad. Ci. Biol. (Granada) 3: 92. 1974 (art. 31) non F. Casas in Ars. Pharm. 11: 274. 1970 (art. 43), Petrocoptidetum crassifoliae O. Bolòs & P. Montserrat ex F. Casas 1970 crassifolietosum G. Montserrat in Lucas Mallada 1: 108. 1989 (corresp. name)]

Typus associatio: Table 69, rel. 1 [Huesca: Lafortunada, Paso de las Devotas, overhang limestone wall. 42º 35'N-0º 10'E. 750 m, W, 20 m2].

Characteristic species: Petrocoptis crassifolia.

Diagnosis: Association well characterized by the endemic Petrocoptis crassifolia, that in the meso-supratemperate submediterranean belt grows in overhanged calcareous walls of the high Cinca river canyons in the huesca Prepyrenees: the Devotas Canyon, Escuaín canyon and Añisclo Valley.

Table 69

29.3.3 Asplenio csikii-Petrocoptidetum crassifoliae
(Valeriano-Petrocoptidion, Petrocoptidetalia, Petrocoptido-Sarcocapnetea)

Altitude (1=10m) 75 73 73 73
Number of species 3 3 3 3
Ordinal number 1* 2 3 4
Characteristic species:
Petrocoptis crassifolia 2 3 2 3
Asplenium csikii 1 1 + 3
Sarcocapnos enneaphylla . 1 . 1
Companion species:
Saxifraga longifolia + . . 1
Adiantum capillus-veneris . . + 1

Localities: 1. Holotypus ass. Huesca: Lafortunada, Paso de las Devotas, overhang limestone wall. 42º 35'N-0º 10'E. W, 20 m2. 2. Huesca: Lafortunada, Paso de las Devotas, overhang limestone wall. E, 10 m2. 3. Huesca: Lafortunada, Paso de las Devotas, moist overhang limestone wall. E, 10 m2. 4. Synthesized table.

[Rivas-Martínez, Costa & P. Soriano] (*)

Asplenio csikii-Petrocoptidetum pseudoviscosae ass. nova hoc loco (29.3.4) (*)

[Petrocoptidetum crassifoliae pseudoviscetosum G. Montserrat in Lucas Mallada 1: 103. 1989 (corresp. name)]

Typus associatio: Table 70, rel. 2 [Huesca: Castejón de Sos, Congosto del Ventamillo, overhang limestone wall. 42º 31'N-0º 28'E. 850 m, W, 10 m2].

Characteristic species: Petrocoptis pseudoviscosa.

Diagnosis: Association well characterized by the local endemic Petrocoptis pseudoviscosa which grows on supratemperate submediterranean limestone and dolomitic limestone overhanged canyons of Ésera river between Campo and Castejón de Sos in Ribagorzian Prepyrenean.

Table 70

29.3.4 Asplenio csikii-Petrocoptidetum pseudoviscosae
(Valeriano-Petrocoptidion, Petrocoptidetalia, Petrocoptido-Sarcocapnetea)

Altitude (1=10m) 83 85 75 75 80
Number of species 4 3 4 5 4
Ordinal number 1 2* 3 4 5
Characteristic species:
Petrocoptis pseudoviscosa 2 2 1 2 4
Asplenium csikii + 1 + 1 4
Sarcocapnos enneaphylla . . + . 1
Companion species:
Saxifraga longifolia + . . + 2
Hieracium phlomoides . + . + 2
Lonicera pyrenaica + . . . 1
Asplenium petrarchae . . + . 1
Asplenium ruta-muraria . . . + 1

Localities: 1. Huesca: Castejón de Sos, Congosto del Ventamillo, overhang limestone wall. W, 20 m2. 2. Holotypus ass. Huesca: Castejón de Sos, Congosto del Ventamillo, overhang limestone wall. W, 10 m2. 42º 31'N-0º 28'E. 3. Huesca: Campo, Ésera canyon, overhang limestone wall. SE, 10 m2. 4. Huesca: Campo, Peña Madrid overhang wall. E, 20 m2. 5. Synthesized table.

[Rivas-Martínez, Costa & P. Soriano] (*)

Petrocoptidetum hispanicae O. Bolòs & P. Montserrat ex F. Casas in Ars. Pharm. 11: 273. 1970 corr. hoc loco (29.3.5)

[Petrocoptidetum crassifoliae O. Bolòs & P. Montserrat ex F. Casas in Ars. Pharm. 11: 273. 1970 (art. 43); excl. réls.: Devotas, Ventamillo and Añisclo]

Distribution: West Pyrenean meso-supratemperate subhumid overhanged calcareous walls on river canyons western to Subordan River.

Petrocoptidetum montserratii ass. nova hoc loco (29.3.6)

Typus associatio: Huesca: Riglos, Mallo Cored, cave and overhanget wall at the bottom of Western face. 42º 20'N-0º 44'W. SW, 820 m, 20 m2. Characteristic species: 3 Petrocoptis montserratii. Companion species: + Adiantum capillus-veneris, + Asplenium ceterach, + Parietaria judaica.

Characteristic species: Petrocoptis montserratii.

Diagnosis: Rupicolous ombrophobous chasmophytic community that grows on shady overhangs and caves of conglomerate rocks in the Mallos de Riglos and Agüero (Huesca). It is also known from other prepyrenaic mountains in Huesca Province such as the Canciás and Santa Orosia.

Petrocoptidetum pyrenaicae F. Casas in Ars. Pharm. 11: 274. 1970 (29.3.7)

[Saxifrago longifoliae-Petrocoptidetum pyrenaicae Rivas-Martínez, Báscones, T.E. Díaz, Fernández-González & Loidi in Itinera Geobot. 5: 381, tb. 67. 1991 (syntax. syn.)]

Distribution: West Pyrenean supra-orotemperate humid-hyperhumid on overhanged calcareous walls.

Petrocoptidetum pardoi F. Casas in Cuad. Ci. Biol. (Granada) 3: 91, tb. 1. 1974 (29.3.8)

Distribution: Supratemperate submediterranean on Maestratian Catalanid Ranges: canyons of Morella-Bergantes river (Santuario de la Virgen de la Balma) on overhanged calcareous walls, characterized by the local endemic Petrocoptis pardoi subsp. pardoi.

Petrocoptido hispanicae-Androsacetum willkommii F. Casas 1970 corr. Benito in Villar & Benito in Mem. Mapa Veg. actual. Parque Nal. Ordesa y Monte Perdido: 139. 2001 (29.3.9)

[Petrocoptido hispanicae-Androsacetum cylindricae F. Casas in Ars. Pharm. 11: 273, tb. 2. 1970 (art. 43)]

Distribution: Prepyrenean Jacetanian (Peña Oroel upper supra-lower orotemperate humid, on north face overhangs).

Petrocoptido montsiccianae-Antirrhinetum mollis O. Bolòs 1954 corr. hoc loco (29.3.10)

[Petrocoptido pardoi-Antirrhinetum mollis O. Bolòs in Collect. Bot. (Barcelona) 4(2): 253, tb. 1. 1954 (art. 43)]

Distribution: Meso-supramediterranean and submediterranean on Montsiccianian canyons of Noguera Pallaresa and Segre: Congosts of Terradets, Collegats and Oliana, characterized by the endemic Petrocoptis pardoi subsp. montsicciana.

Pinguiculo longifoliae-Androsacetum cylindricae F. Casas in Ars. Pharm. 11: 278. 1970 (29.3.11)

Distribution: Central Pyrenean orotemperate humid on overhanged calcareous wet walls from Ordesa National Park to Cotiella Peak.

Saxifrago longifoliae-Valerianetum longiflorae Rivas Goday in Rivas Goday, Esteve, Rigual & Borja in Anales Inst. Bot. Cavanilles 12(1): 487, tb. 3. 1954 (29.3.12)

Distribution: Prepyrenean Guarensian supratemperate subhumid submediterranean, described of Arguis Mountains (Guara Mountains), characterized by Valeriana longiflora subsp. longiflora. It seems intermediate with Saxifrago longifoliae-Ramondetum myconi.

Scrophulario pyrenaicae-Antirrhinetum sempervirentis Quézel in Collect. Bot. (Barcelona) 5(1): 185, tb. 5. 1956 (29.3.13)

[Scrophularietum pyrenaicae F. Casas in Trab. Dep. Bot. Univ. Granada Dep. Bot. Univ. Granada 1: 32. 1970]

Distribution: Central Pyrenean supra-orotemperate humid on overhanged calcareous cliff and crevices visited by birds from Ordesa National Park to Cotiella Peak.

Valeriano longiflorae-Petrocoptidetum guarensis F. Casas 1970 corr. hoc loco (29.3.14)

[Valeriano longiflorae-Petrocoptidetum montsiccianae F. Casas in Ars. Pharm. 11: 274, tb. 3. 1970 (art. 43)]

Distribution: Meso-supramediterranean and submediterranean in Jacetanian-Guarensian canyons like Mascún canyon of the basin of Alcanadre river in Sierra de Guara range, characterized by the endemic Petrocoptis pardoi subsp. guarensis.

Valeriano longiflorae-Petrocoptidetum hispanicae O. Bolòs & P. Montserrat ex F. Casas in Ars. Pharm. 11: 274. 1970 (29.3.15)

Distribution: Prepyrenean Jacetanian supratemperate submediterranean subhumid (San Juan de la Peña), characterized by Petrocoptis pyrenaica subsp. hispanica.

Petrocoptidion glaucifoliae (P. Fernández, Penas & T.E. Díaz 1983) all. nova, stat. nov. hoc loco (29.4)

[Petrocoptidenion glaucifoliae P. Fernández, Penas & T.E. Díaz in Anales Jardín Bot. Madrid 40 (1): 220. 1983 (art. 27a), Petrocoptidion cantabricum F. Casas in Trab. Dep. Bot. Univ. Granada 1: 31. 1972 (art. 2b, 8), holotypus: Petrocoptidetum cantabricum Rothmaler in Bot. Jahrb. 72: 119. 1941 (art. 2b, 7), Rupicampanulion Rothmaler in Vegetatio 5-6: 599. 1954 (art. 2b), Petrocoptidenion grandifloro-viscosae P. Fernández, Penas & T.E. Díaz in Anales Jard. Bot. Madrid 40(1): 226. 1983 (corresp. name)]

Typus alliancia: Petrocoptidetum glaucifoliae Rivas-Martínez in P. Fernández, Penas & T.E. Díaz in Anales Jard. Bot. Madrid 40(1): 220. 1983.

Characteristic species: Petrocoptis pyrenaica subsp. glaucifolia (29.4), Petrocoptis grandiflora (29.4), Petrocoptis pyrenaica subsp. wiedmannii (29.4), Petrocoptis pyrenaica subsp. viscosa (29.4).

Diagnosis: Supra-oromediterranean and meso-supramediterranean overhanging dolomite and limestone communities in Orocantabric and Bercian mountain.

Petrocoptidetum glaucifoliae Rivas-Martínez in P. Fernández, Penas & T.E. Díaz in Anales Jard. Bot. Madrid 40(1): 226, tb. 4. 1983 (29.4.1)

Distribution: Ubinnean-Picoeuropean supra to upperorotemperate humid and hyperhumid overhanging limestone community, characterized by Petrocoptis pyrenaica subsp. glaucifolia.

Petrocoptidetum grandiflorae J.M. Losa, Mayor, Andrés & F. Navarro in Anales Inst. Bot. Cavanilles 32(1): 215, tb. 2. 1975 (29.4.2)

Distribution: Bercian meso-supramediterranean subhumid growing on dolomitic-limestone overhanged walls, confined to calcareous and dolostone mountains cut by Sil River near Carucedo, characterized by Petrocoptis grandiflora.

Petrocoptidetum viscosae J.M. Losa, Mayor, Andrés & F. Navarro in Anales Inst. Bot. Cavanilles 32(1): 215, tb. 1. 1975 (29.4.3)

Distribution: Bercian mesomediterranean subhumid growing on dolomitic-limestone walls, overhanged only on primary stations, confined to Priaranza del Bierzo and Cornatel Castle dolostone zone, characterized by Petrocoptis pyrenaica subsp. viscosa.

Petrocoptidetum wiedmannii Ladero, T.E. Díaz, Penas, Rivas-Martínez & C. Valle in Itinera Geobot. 1: 70. 1987 (29.4.4)

Distribution: Ubinnean-Picoeuropean and Ovetensean meso-lowersupratemperate humid, growing on calcareous overhang walls, characterized by Petrocoptis pyrenaica subsp. wiedmannii.

[Rivas-Martínez, Cantó & Izco]